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A fruiting fig tree outside of our cabins at Monteverde attracted a variety of hungry birds, including this lovely and cooperative Emerald Toucanet, photographed by tour participant Chris DeCilio
Without a doubt, this year's classic Costa Rica was by far the driest ever. Even in the dry season, a certain amount of rain is expected, but the minimal amount of rain we experienced on this trip was low, even by dry season standards. Fortunately, though much of the country was pretty sere, there was nothing dry about the birding, and the dry weather meant that we didn't lose any valuable birding time to rain!
Our birding got off to a fine start the first morning when we spotted some raptors circling around in the clear morning sky near Virgen del Socorro. Hopping off the bus, we were treated to eye level views of beautiful Swallow-tailed Kites and White Hawks rising up on the mid-morning thermals. Before that first day was complete, we'd been buzzed by hordes of hummers at the La Paz Waterfall Gardens, notched our first trogon and toucan, and come face to face (almost) with a roosting Great Potoo! All in all it wasn't a bad introduction to what CR has to offer!
The next few days were full of highlights as we birded the rich Caribbean lowlands in the La Selva region. Along the wonderful entrance road to the OTS station, we enjoyed a parade of great birds: nearly side-by-side views of Rufous and Broad-billed motmots; a lovely pair of Cinnamon Woodpeckers intently working over a roadside tree trunk just a few meters away; a pair of Slaty-tailed Trogons excavating a nest cavity in an arboreal termitary; immaculate Snowy Cotingas chasing each other around above the canopy; a pair of Rufous Mourners landing in the road ahead of us. The reserve itself was a bit quieter, but still provided some memorable moments, such as the male Great Curassow singing from a high perch near the trail and that responsive Vermiculated Screech-Owl that nearly landed on Lenin's head! Elsewhere in the area, both Scarlet and Great Green macaws offered exceptional views, and the sight of hundreds of Turkey Vultures and Swainson's Hawks streaming northward above us was simply amazing.
The cooler temperatures at Rancho Naturalista were most welcome after the heat of the lowlands, and a different mix of birds awaited us here, too. A low-soaring Black Hawk-Eagle flying across a wooded hillside, a pair of stunning Sunbitterns along a clear mountain stream, and the parade of birds at the "moth cloth" -- Buff-throated Foliage-gleaner, the very local Tawny-chested Flycatcher, Stripe-breasted Wren, and Red-throated Ant-Tanager, among others -- were a few of the many birds that kept things exciting during our time here. Our final Caribbean slope birding came at Tapanti National Park as we moved on up into the mountains. A bold Streak-breasted Treehunter, an excited pair of Black-and-white Becards, a striking male Red-headed Barbet, and brilliant Golden-browed Chlorophonias made our short visit there exceedingly sweet!
Chiriqui specialties were the theme in the highlands of the Cerro de la Muerte region, where many species restricted to Panama and CR can be found. We encountered more than 30 of these Chiriqui endemics here, a list that included such treats as the scarce Silvery-throated Jay that suddenly appeared in the forest beside us, the skulking Wrenthrush in the thick bamboo along the Savegre River, the charming Collared Redstarts that flitted about on the trailsides, and the glittering Fiery-throated Hummingbirds that darted about at the feeders of La Georgina, their fiery throats glowing brilliantly when the angle of the light was just right. Despite all the local endemics, it was actually a much more widespread bird that stole the show: the Resplendent Quetzal! Having a pair of these showy birds nesting right behind our cabins was a real thrill!
Dropping to the Pacific coast, we were back in the heat and among another completely different suite of birds. The forests of the Carara region gifted us a bunch of superb birds: a Crested Owl growling in the canopy of a roadside tree; Golden-naped Woodpecker and Baird's Trogon, two local specialties, seen beautifully along the Quebrada Bonita trail; a couple of incredibly confiding Orange-collared Manakins at their trailside lek; a surprise Streak-chested Antpitta hopping along the trail ahead of us. Away from the forest, such species as Roseate Spoonbill, Double-striped Thick-knee, Gray Hawk, Striped Owl, and Turquoise-browed Motmot added to the birding excitement.
We finished up in the beautiful cloud forests of Monteverde. Foremost among our targets was the bizarre and spectacular Three-wattled Bellbird, which gave a brief but memorable performance from a dead snag nearly right overhead. I think my ears are still ringing! Black-breasted Wood-Quail, dainty Magenta-throated Woodstars, White-fronted Parrots, a couple of gorgeous male Long-tailed Manakins, Ruddy Woodcreeper (at an army ant swarm) and White-naped Brush-Finches were among the many other species we encountered here.
This tour also featured a pretty good list of both mammals and herps, many of which we seldom see on our trips (and a few that were "lifers" for your guides, too). Among the mammals, the sleepy Mexican Hairy Porcupine beside the trail at La Paz, the super Northern Tamandua at Braulio Carrillo, the pair of Vesper Rats (a first for me) on the La Selva night walk, and the curious Long-tailed Weasel at Savegre stand out. As for the herps, well that incredible large Neotropical Chameleon and the Yellow-spotted Night Lizard in its trailside burrow were both first for me!
Finally, I just have to say that, as great as the birding was, the big highlight for Megan and me was having such a great group with whom to share them all. We truly enjoyed your camaraderie and companionship, and we look forward to traveling with you all again someday soon. Many thanks for joining us on the trip. Thanks, too, to our driver, Lenin, for getting us around the country safely and professionally. Happy birding!
--Jay
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Tinamidae (Tinamous)

Large game birds are scarce where hunting is permitted, but they can become quite habituated where they are undisturbed by people, like this Crested Guan at La Selva. (Photo by guide Megan Crewe)
GREAT TINAMOU (Tinamus major) [*] LITTLE TINAMOU (Crypturellus soui) [*]
Anatidae (Ducks, Geese, and Waterfowl)
BLACK-BELLIED WHISTLING-DUCK (Dendrocygna autumnalis)
MUSCOVY DUCK (Cairina moschata)
BLUE-WINGED TEAL (Anas discors) [b]
RING-NECKED DUCK (Aythya collaris) [b]
LESSER SCAUP (Aythya affinis) [b]
Cracidae (Guans, Chachalacas, and Curassows)
GRAY-HEADED CHACHALACA (Ortalis cinereiceps)
CRESTED GUAN (Penelope purpurascens)
BLACK GUAN (Chamaepetes unicolor)
GREAT CURASSOW (Crax rubra)
Odontophoridae (New World Quail)
BUFFY-CROWNED WOOD-PARTRIDGE (Dendrortyx leucophrys) [*]
BLACK-BREASTED WOOD-QUAIL (Odontophorus leucolaemus)
SPOTTED WOOD-QUAIL (Odontophorus guttatus)
Podicipedidae (Grebes)
LEAST GREBE (Tachybaptus dominicus)
Ciconiidae (Storks)
WOOD STORK (Mycteria americana)
Fregatidae (Frigatebirds)
MAGNIFICENT FRIGATEBIRD (Fregata magnificens)
Phalacrocoracidae (Cormorants and Shags)

Arguably the most responsive bird of the trip was this Vermiculated Screech-Owl, which nearly hit Lenin in the head as it flew in, then perched right beside the trail and started singing as we watch from a few feet away. Amazing! (Photo by guide Jay VanderGaast)
NEOTROPIC CORMORANT (Phalacrocorax brasilianus) Anhingidae (Anhingas)
ANHINGA (Anhinga anhinga)
Pelecanidae (Pelicans)
BROWN PELICAN (Pelecanus occidentalis)
Ardeidae (Herons, Egrets, and Bitterns)
BARE-THROATED TIGER-HERON (Tigrisoma mexicanum)
GREAT BLUE HERON (Ardea herodias) [b]
GREAT EGRET (Ardea alba)
SNOWY EGRET (Egretta thula)
LITTLE BLUE HERON (Egretta caerulea)
TRICOLORED HERON (Egretta tricolor)
CATTLE EGRET (Bubulcus ibis)
GREEN HERON (Butorides virescens)
BLACK-CROWNED NIGHT-HERON (Nycticorax nycticorax)
YELLOW-CROWNED NIGHT-HERON (Nyctanassa violacea)
Threskiornithidae (Ibises and Spoonbills)
WHITE IBIS (Eudocimus albus)
GREEN IBIS (Mesembrinibis cayennensis)
ROSEATE SPOONBILL (Platalea ajaja)
Cathartidae (New World Vultures)
BLACK VULTURE (Coragyps atratus)
TURKEY VULTURE (Cathartes aura)
KING VULTURE (Sarcoramphus papa)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus) [b]
Accipitridae (Hawks, Eagles, and Kites)
WHITE-TAILED KITE (Elanus leucurus)
HOOK-BILLED KITE (Chondrohierax uncinatus)
GRAY-HEADED KITE (Leptodon cayanensis)
SWALLOW-TAILED KITE (Elanoides forficatus)

Swallow-tailed Kites were a common sight on the tour. This was one of several that were soaring with White Hawks, Barred Hawks, and a Broad-winged Hawk over the beautiful Virgen del Socorro valley on our first morning. (Photo by tour participant Marshall Dahl)
BLACK HAWK-EAGLE (Spizaetus tyrannus) ORNATE HAWK-EAGLE (Spizaetus ornatus) [*]
SNAIL KITE (Rostrhamus sociabilis)
PLUMBEOUS KITE (Ictinia plumbea)
COMMON BLACK-HAWK (MANGROVE) (Buteogallus anthracinus subtilis)
BARRED HAWK (Morphnarchus princeps)
ROADSIDE HAWK (Rupornis magnirostris)
WHITE HAWK (Pseudastur albicollis)
BROAD-WINGED HAWK (Buteo platypterus) [b]
GRAY HAWK (Buteo plagiatus)
SHORT-TAILED HAWK (Buteo brachyurus)
SWAINSON'S HAWK (Buteo swainsoni) [b]
RED-TAILED HAWK (Buteo jamaicensis)
Eurypygidae (Sunbittern)
SUNBITTERN (Eurypyga helias)
Rallidae (Rails, Gallinules, and Coots)
WHITE-THROATED CRAKE (Laterallus albigularis) [*]
GRAY-NECKED WOOD-RAIL (Aramides cajaneus)
PURPLE GALLINULE (Porphyrio martinicus)
Aramidae (Limpkin)
LIMPKIN (Aramus guarauna)
Burhinidae (Thick-knees)
DOUBLE-STRIPED THICK-KNEE (Burhinus bistriatus)
Recurvirostridae (Stilts and Avocets)

Tour participant Pete Thayer spotted (and photographed) this Sunbittern along the river at La Mina. We were fortunate to also get to see the stunning wing pattern of this bird as it flew a short way upriver a couple of times.
BLACK-NECKED STILT (Himantopus mexicanus) Charadriidae (Plovers and Lapwings)
SOUTHERN LAPWING (Vanellus chilensis)
KILLDEER (Charadrius vociferus) [b]
Jacanidae (Jacanas)
NORTHERN JACANA (Jacana spinosa)
Scolopacidae (Sandpipers and Allies)
SPOTTED SANDPIPER (Actitis macularius) [b]
SOLITARY SANDPIPER (Tringa solitaria) [b]
GREATER YELLOWLEGS (Tringa melanoleuca) [b]
WILLET (Tringa semipalmata) [b]
WHIMBREL (Numenius phaeopus) [b]
LEAST SANDPIPER (Calidris minutilla) [b]
Laridae (Gulls, Terns, and Skimmers)
LAUGHING GULL (Leucophaeus atricilla) [b]
ROYAL TERN (Thalasseus maximus)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
PALE-VENTED PIGEON (Patagioenas cayennensis)
RED-BILLED PIGEON (Patagioenas flavirostris)
BAND-TAILED PIGEON (Patagioenas fasciata)
RUDDY PIGEON (Patagioenas subvinacea)
SHORT-BILLED PIGEON (Patagioenas nigrirostris)

We were very pleased to find this Black-and-white Owl back on its day roost at Orotina park, as it had been missing here for some time beforehand. (Photo by guide Megan Crewe)
WHITE-WINGED DOVE (Zenaida asiatica) INCA DOVE (Columbina inca)
COMMON GROUND-DOVE (Columbina passerina)
RUDDY GROUND-DOVE (Columbina talpacoti)
WHITE-TIPPED DOVE (Leptotila verreauxi)
GRAY-CHESTED DOVE (Leptotila cassini)
Cuculidae (Cuckoos)
SQUIRREL CUCKOO (Piaya cayana)
LESSER GROUND-CUCKOO (Morococcyx erythropygus)
SMOOTH-BILLED ANI (Crotophaga ani)
GROOVE-BILLED ANI (Crotophaga sulcirostris)
Strigidae (Owls)
TROPICAL SCREECH-OWL (Megascops choliba)
VERMICULATED SCREECH-OWL (GUATEMALAN) (Megascops guatemalae vermiculatus)
BARE-SHANKED SCREECH-OWL (Megascops clarkii)
CRESTED OWL (Lophostrix cristata)
FERRUGINOUS PYGMY-OWL (Glaucidium brasilianum)
MOTTLED OWL (Ciccaba virgata)
BLACK-AND-WHITE OWL (Ciccaba nigrolineata)
STRIPED OWL (Pseudoscops clamator)
Caprimulgidae (Nightjars and Allies)
LESSER NIGHTHAWK (Chordeiles acutipennis)
SHORT-TAILED NIGHTHAWK (Lurocalis semitorquatus)
COMMON PAURAQUE (Nyctidromus albicollis)
DUSKY NIGHTJAR (Antrostomus saturatus)
Nyctibiidae (Potoos)
GREAT POTOO (Nyctibius grandis)
Apodidae (Swifts)

Costa Rica is always a good place for hummers. This White-necked Jacobin was one of more than 35 species we saw on the trip. (Photo by tour participant Chris DeCilio)
CHESTNUT-COLLARED SWIFT (Streptoprocne rutila) WHITE-COLLARED SWIFT (Streptoprocne zonaris)
VAUX'S SWIFT (Chaetura vauxi)
GRAY-RUMPED SWIFT (Chaetura cinereiventris)
LESSER SWALLOW-TAILED SWIFT (Panyptila cayennensis)
Trochilidae (Hummingbirds)
WHITE-NECKED JACOBIN (Florisuga mellivora)
BAND-TAILED BARBTHROAT (Threnetes ruckeri)
GREEN HERMIT (Phaethornis guy)
LONG-BILLED HERMIT (Phaethornis longirostris)
STRIPE-THROATED HERMIT (Phaethornis striigularis)
GREEN-FRONTED LANCEBILL (Doryfera ludovicae)
BROWN VIOLETEAR (Colibri delphinae)
GREEN VIOLETEAR (Colibri thalassinus)
PURPLE-CROWNED FAIRY (Heliothryx barroti)
GREEN-BREASTED MANGO (Anthracothorax prevostii)
GREEN THORNTAIL (Discosura conversii)
GREEN-CROWNED BRILLIANT (Heliodoxa jacula)
MAGNIFICENT HUMMINGBIRD (Eugenes fulgens)
PLAIN-CAPPED STARTHROAT (Heliomaster constantii)
FIERY-THROATED HUMMINGBIRD (Panterpe insignis)
WHITE-BELLIED MOUNTAIN-GEM (Lampornis hemileucus)
PURPLE-THROATED MOUNTAIN-GEM (Lampornis calolaemus)
WHITE-THROATED MOUNTAIN-GEM (GRAY-TAILED) (Lampornis castaneoventris cinereicauda)
MAGENTA-THROATED WOODSTAR (Calliphlox bryantae)
VOLCANO HUMMINGBIRD (Selasphorus flammula)
SCINTILLANT HUMMINGBIRD (Selasphorus scintilla)
VIOLET-HEADED HUMMINGBIRD (Klais guimeti)
SCALY-BREASTED HUMMINGBIRD (Phaeochroa cuvierii)
VIOLET SABREWING (Campylopterus hemileucurus)

One of the prizes of our visit to the La Paz Waterfall Gardens was this Black-bellied Hummingbird, which is restricted to the mountains of Costa Rica and Panama. (Photo by guide Megan Crewe)
BRONZE-TAILED PLUMELETEER (Chalybura urochrysia) CROWNED WOODNYMPH (Thalurania colombica)
STRIPE-TAILED HUMMINGBIRD (Eupherusa eximia)
BLACK-BELLIED HUMMINGBIRD (Eupherusa nigriventris)
WHITE-TAILED EMERALD (Elvira chionura)
COPPERY-HEADED EMERALD (Elvira cupreiceps) [E]
SNOWCAP (Microchera albocoronata)
BLUE-CHESTED HUMMINGBIRD (Amazilia amabilis)
MANGROVE HUMMINGBIRD (Amazilia boucardi) [E]
STEELY-VENTED HUMMINGBIRD (Amazilia saucerrottei)
SNOWY-BELLIED HUMMINGBIRD (Amazilia edward)
RUFOUS-TAILED HUMMINGBIRD (Amazilia tzacatl)
CINNAMON HUMMINGBIRD (Amazilia rutila)
Trogonidae (Trogons)
RESPLENDENT QUETZAL (Pharomachrus mocinno)
SLATY-TAILED TROGON (Trogon massena)
BLACK-HEADED TROGON (Trogon melanocephalus)
BAIRD'S TROGON (Trogon bairdii)

A male Slaty-tailed Trogon takes a pause from excavating a nest in an arboreal termitary at La Selva. (Photo by guide Jay VanderGaast)
GARTERED TROGON (Trogon caligatus) BLACK-THROATED TROGON (Trogon rufus)
ORANGE-BELLIED TROGON (Trogon aurantiiventris) [*]
COLLARED TROGON (Trogon collaris)
Momotidae (Motmots)
BLUE-CROWNED MOTMOT (LESSON'S) (Momotus coeruliceps lessonii)
RUFOUS MOTMOT (Baryphthengus martii)
BROAD-BILLED MOTMOT (Electron platyrhynchum)
TURQUOISE-BROWED MOTMOT (Eumomota superciliosa)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)
BELTED KINGFISHER (Megaceryle alcyon) [b]
AMAZON KINGFISHER (Chloroceryle amazona)
GREEN KINGFISHER (Chloroceryle americana)
AMERICAN PYGMY KINGFISHER (Chloroceryle aenea)
Bucconidae (Puffbirds)
WHITE-NECKED PUFFBIRD (Notharchus hyperrhynchus)
PIED PUFFBIRD (Notharchus tectus)
WHITE-WHISKERED PUFFBIRD (Malacoptila panamensis)
Galbulidae (Jacamars)
RUFOUS-TAILED JACAMAR (Galbula ruficauda)
Capitonidae (New World Barbets)
RED-HEADED BARBET (Eubucco bourcierii)
Semnornithidae (Toucan-Barbets)
PRONG-BILLED BARBET (Semnornis frantzii)
Ramphastidae (Toucans)
EMERALD TOUCANET (BLUE-THROATED) (Aulacorhynchus prasinus caeruleogularis)
COLLARED ARACARI (Pteroglossus torquatus)
BLACK-MANDIBLED TOUCAN (CHESTNUT-MANDIBLED) (Ramphastos ambiguus swainsonii)

White-whiskered Puffbirds are rather quiet and inconspicuous, so it was great to find this one sitting quietly near the trail at Carara. (Photo by tour participant Pete Thayer)
KEEL-BILLED TOUCAN (Ramphastos sulfuratus) Picidae (Woodpeckers)
ACORN WOODPECKER (Melanerpes formicivorus)
GOLDEN-NAPED WOODPECKER (Melanerpes chrysauchen)
BLACK-CHEEKED WOODPECKER (Melanerpes pucherani)
RED-CROWNED WOODPECKER (Melanerpes rubricapillus)
HOFFMANN'S WOODPECKER (Melanerpes hoffmannii)
SMOKY-BROWN WOODPECKER (Picoides fumigatus) [*]
HAIRY WOODPECKER (Picoides villosus) [*]
RUFOUS-WINGED WOODPECKER (Piculus simplex)
GOLDEN-OLIVE WOODPECKER (Colaptes rubiginosus)
CINNAMON WOODPECKER (Celeus loricatus)
LINEATED WOODPECKER (Dryocopus lineatus)
Falconidae (Falcons and Caracaras)
COLLARED FOREST-FALCON (Micrastur semitorquatus) [*]
CRESTED CARACARA (Caracara cheriway)
YELLOW-HEADED CARACARA (Milvago chimachima)
LAUGHING FALCON (Herpetotheres cachinnans) [*]
BAT FALCON (Falco rufigularis)
PEREGRINE FALCON (Falco peregrinus) [b]
Psittacidae (Parrots)
SULPHUR-WINGED PARAKEET (Pyrrhura hoffmanni)
CRIMSON-FRONTED PARAKEET (Aratinga finschi)
OLIVE-THROATED PARAKEET (AZTEC) (Aratinga nana astec)
ORANGE-FRONTED PARAKEET (Aratinga canicularis)
GREAT GREEN MACAW (Ara ambiguus)
SCARLET MACAW (Ara macao)
BARRED PARAKEET (Bolborhynchus lineola)
ORANGE-CHINNED PARAKEET (Brotogeris jugularis)
BROWN-HOODED PARROT (Pyrilia haematotis)
WHITE-CROWNED PARROT (Pionus senilis)

A pair of Cinnamon Woodpeckers along the La Selva entrance road gave a wonderful show as they worked over a tree trunk low down next to the road. This one is the male. (Photo by tour participant Chris De Cilio)
RED-LORED PARROT (Amazona autumnalis) YELLOW-NAPED PARROT (Amazona auropalliata)
WHITE-FRONTED PARROT (Amazona albifrons)
Thamnophilidae (Typical Antbirds)
FASCIATED ANTSHRIKE (Cymbilaimus lineatus)
BARRED ANTSHRIKE (Thamnophilus doliatus)
BLACK-CROWNED ANTSHRIKE (Thamnophilus atrinucha)
BLACK-HOODED ANTSHRIKE (Thamnophilus bridgesi)
RUSSET ANTSHRIKE (Thamnistes anabatinus)
PLAIN ANTVIREO (Dysithamnus mentalis)
STREAK-CROWNED ANTVIREO (Dysithamnus striaticeps)
CHECKER-THROATED ANTWREN (Epinecrophylla fulviventris)
SLATY ANTWREN (Myrmotherula schisticolor) [*]
DOT-WINGED ANTWREN (Microrhopias quixensis)
DUSKY ANTBIRD (Cercomacra tyrannina)
CHESTNUT-BACKED ANTBIRD (Myrmeciza exsul)
DULL-MANTLED ANTBIRD (Myrmeciza laemosticta) [*]
ZELEDON'S ANTBIRD (Myrmeciza zeledoni) [*]
Grallariidae (Antpittas)
STREAK-CHESTED ANTPITTA (Hylopezus perspicillatus)
THICKET ANTPITTA (Hylopezus dives) [*]
Rhinocryptidae (Tapaculos)
SILVERY-FRONTED TAPACULO (Scytalopus argentifrons)
Furnariidae (Ovenbirds and Woodcreepers)
TAWNY-THROATED LEAFTOSSER (Sclerurus mexicanus)
OLIVACEOUS WOODCREEPER (Sittasomus griseicapillus)
PLAIN-BROWN WOODCREEPER (Dendrocincla fuliginosa)
RUDDY WOODCREEPER (Dendrocincla homochroa)
WEDGE-BILLED WOODCREEPER (Glyphorynchus spirurus)
NORTHERN BARRED-WOODCREEPER (Dendrocolaptes sanctithomae)
COCOA WOODCREEPER (Xiphorhynchus susurrans)

Streak-breasted Treehunters don't usually pose out in the open like this, but this one at Tapanti was just so curious. (Photo by tour participant Marshall Dahl)
BLACK-STRIPED WOODCREEPER (Xiphorhynchus lachrymosus) [*] SPOTTED WOODCREEPER (Xiphorhynchus erythropygius)
BROWN-BILLED SCYTHEBILL (Campylorhamphus pusillus) [*]
STREAK-HEADED WOODCREEPER (Lepidocolaptes souleyetii)
SPOT-CROWNED WOODCREEPER (Lepidocolaptes affinis)
PLAIN XENOPS (Xenops minutus)
BUFFY TUFTEDCHEEK (Pseudocolaptes lawrencii)
LINEATED FOLIAGE-GLEANER (Syndactyla subalaris)
STRIPED WOODHAUNTER (Hyloctistes subulatus) [*]
BUFF-THROATED FOLIAGE-GLEANER (HYPOPHAEUS) (Automolus ochrolaemus hypophaeus)
STREAK-BREASTED TREEHUNTER (Thripadectes rufobrunneus)
SPOTTED BARBTAIL (Premnoplex brunnescens)
RUDDY TREERUNNER (Margarornis rubiginosus)
RED-FACED SPINETAIL (Cranioleuca erythrops)
Tyrannidae (Tyrant Flycatchers)
NORTHERN BEARDLESS-TYRANNULET (Camptostoma imberbe)
SOUTHERN BEARDLESS-TYRANNULET (Camptostoma obsoletum)
YELLOW TYRANNULET (Capsiempis flaveola)
GREENISH ELAENIA (Myiopagis viridicata)
YELLOW-BELLIED ELAENIA (Elaenia flavogaster)
MOUNTAIN ELAENIA (Elaenia frantzii)
TORRENT TYRANNULET (Serpophaga cinerea)
OLIVE-STRIPED FLYCATCHER (Mionectes olivaceus)
OCHRE-BELLIED FLYCATCHER (Mionectes oleagineus)
SLATY-CAPPED FLYCATCHER (Leptopogon superciliaris)
PALTRY TYRANNULET (Zimmerius vilissimus)
BLACK-CAPPED PYGMY-TYRANT (Myiornis atricapillus) [*]
SCALE-CRESTED PYGMY-TYRANT (Lophotriccus pileatus)
NORTHERN BENTBILL (Oncostoma cinereigulare)
SLATE-HEADED TODY-FLYCATCHER (Poecilotriccus sylvia)
COMMON TODY-FLYCATCHER (Todirostrum cinereum)
BLACK-HEADED TODY-FLYCATCHER (Todirostrum nigriceps)
EYE-RINGED FLATBILL (Rhynchocyclus brevirostris)
YELLOW-OLIVE FLYCATCHER (Tolmomyias sulphurescens)
WHITE-THROATED SPADEBILL (Platyrinchus mystaceus)
GOLDEN-CROWNED SPADEBILL (Platyrinchus coronatus)
RUDDY-TAILED FLYCATCHER (Terenotriccus erythrurus)
SULPHUR-RUMPED FLYCATCHER (Myiobius sulphureipygius aureatus)
TAWNY-CHESTED FLYCATCHER (Aphanotriccus capitalis)
TUFTED FLYCATCHER (Mitrephanes phaeocercus)
DARK PEWEE (Contopus lugubris)

This Orange-collared Manakin sat calmly on his display perch next to the trail at Carara, obviously quite accustomed to having people stop by and gawk! (Photo by tour participant Pete Thayer)
EASTERN WOOD-PEWEE (Contopus virens) [b] TROPICAL PEWEE (Contopus cinereus)
YELLOW-BELLIED FLYCATCHER (Empidonax flaviventris) [b]
YELLOWISH FLYCATCHER (Empidonax flavescens)
BLACK-CAPPED FLYCATCHER (Empidonax atriceps)
BLACK PHOEBE (Sayornis nigricans)
LONG-TAILED TYRANT (Colonia colonus)
BRIGHT-RUMPED ATTILA (Attila spadiceus)
RUFOUS MOURNER (Rhytipterna holerythra)
DUSKY-CAPPED FLYCATCHER (Myiarchus tuberculifer)
PANAMA FLYCATCHER (Myiarchus panamensis)
GREAT CRESTED FLYCATCHER (Myiarchus crinitus) [b]
BROWN-CRESTED FLYCATCHER (Myiarchus tyrannulus)
GREAT KISKADEE (Pitangus sulphuratus)
BOAT-BILLED FLYCATCHER (Megarynchus pitangua)
SOCIAL FLYCATCHER (Myiozetetes similis)
GRAY-CAPPED FLYCATCHER (Myiozetetes granadensis)
WHITE-RINGED FLYCATCHER (Conopias albovittatus)
GOLDEN-BELLIED FLYCATCHER (Myiodynastes hemichrysus)
STREAKED FLYCATCHER (Myiodynastes maculatus)
SULPHUR-BELLIED FLYCATCHER (Myiodynastes luteiventris) [a]
PIRATIC FLYCATCHER (Legatus leucophaius) [a]
TROPICAL KINGBIRD (Tyrannus melancholicus)
SCISSOR-TAILED FLYCATCHER (Tyrannus forficatus) [b]
Cotingidae (Cotingas)
PURPLE-THROATED FRUITCROW (Querula purpurata)
RUFOUS PIHA (Lipaugus unirufus) [*]
THREE-WATTLED BELLBIRD (Procnias tricarunculatus)
SNOWY COTINGA (Carpodectes nitidus)
Pipridae (Manakins)
WHITE-RUFFED MANAKIN (Corapipo altera)
LONG-TAILED MANAKIN (Chiroxiphia linearis)
WHITE-CROWNED MANAKIN (Dixiphia pipra)
RED-CAPPED MANAKIN (Ceratopipra mentalis)
WHITE-COLLARED MANAKIN (Manacus candei)
ORANGE-COLLARED MANAKIN (Manacus aurantiacus)
Tityridae (Tityras and Allies)
BLACK-CROWNED TITYRA (Tityra inquisitor)

Timberline Wrens came easy this trip, though that isn't always the case; this one was hopping about in the open scolding when we came walking down Providencia Road. (Photo by tour participant Marshall Dahl)
MASKED TITYRA (Tityra semifasciata) NORTHERN SCHIFFORNIS (Schiffornis veraepacis) [*]
CINNAMON BECARD (Pachyramphus cinnamomeus)
WHITE-WINGED BECARD (Pachyramphus polychopterus)
BLACK-AND-WHITE BECARD (Pachyramphus albogriseus)
ROSE-THROATED BECARD (Pachyramphus aglaiae)
Vireonidae (Vireos)
MANGROVE VIREO (Vireo pallens) [*]
YELLOW-THROATED VIREO (Vireo flavifrons) [b]
YELLOW-WINGED VIREO (Vireo carmioli)
BROWN-CAPPED VIREO (Vireo leucophrys)
PHILADELPHIA VIREO (Vireo philadelphicus) [b]
RED-EYED VIREO (Vireo olivaceus) [b]
YELLOW-GREEN VIREO (Vireo flavoviridis) [a]
TAWNY-CROWNED GREENLET (Hylophilus ochraceiceps)
LESSER GREENLET (Hylophilus decurtatus)
RUFOUS-BROWED PEPPERSHRIKE (Cyclarhis gujanensis) [*]
Corvidae (Crows, Jays, and Magpies)
SILVERY-THROATED JAY (Cyanolyca argentigula)
WHITE-THROATED MAGPIE-JAY (Calocitta formosa)
BROWN JAY (Psilorhinus morio)
Hirundinidae (Swallows)
BLUE-AND-WHITE SWALLOW (Pygochelidon cyanoleuca)
NORTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx serripennis)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)
GRAY-BREASTED MARTIN (Progne chalybea)
MANGROVE SWALLOW (Tachycineta albilinea)
BARN SWALLOW (Hirundo rustica) [b]
Troglodytidae (Wrens)
NIGHTINGALE WREN (Microcerculus philomela) [*]
SCALY-BREASTED WREN (WHISTLING) (Microcerculus marginatus luscinia) [*]
HOUSE WREN (Troglodytes aedon)
OCHRACEOUS WREN (Troglodytes ochraceus)
TIMBERLINE WREN (Thryorchilus browni)
BAND-BACKED WREN (Campylorhynchus zonatus)
RUFOUS-NAPED WREN (Campylorhynchus rufinucha)
BLACK-BELLIED WREN (Pheugopedius fasciatoventris)
RUFOUS-BREASTED WREN (Pheugopedius rutilus)
BLACK-THROATED WREN (Pheugopedius atrogularis) [*]
RUFOUS-AND-WHITE WREN (Thryophilus rufalbus)
STRIPE-BREASTED WREN (Cantorchilus thoracicus)
PLAIN WREN (Cantorchilus modestus)
PLAIN WREN (CANEBRAKE) (Cantorchilus modestus zeledoni)
RIVERSIDE WREN (Cantorchilus semibadius)
BAY WREN (Cantorchilus nigricapillus)
WHITE-BREASTED WOOD-WREN (Henicorhina leucosticta)

Our first Black-cowled Oriole at La Paz was notable not just because it was an unusual record at that high an elevation, but also because it was Marshall's 3000th life bird! (Photo by guide Megan Crewe)
GRAY-BREASTED WOOD-WREN (Henicorhina leucophrys) Polioptilidae (Gnatcatchers)
LONG-BILLED GNATWREN (Ramphocaenus melanurus)
WHITE-LORED GNATCATCHER (Polioptila albiloris)
TROPICAL GNATCATCHER (Polioptila plumbea)
Cinclidae (Dippers)
AMERICAN DIPPER (Cinclus mexicanus)
Turdidae (Thrushes and Allies)
BLACK-FACED SOLITAIRE (Myadestes melanops)
BLACK-BILLED NIGHTINGALE-THRUSH (Catharus gracilirostris)
ORANGE-BILLED NIGHTINGALE-THRUSH (Catharus aurantiirostris)
SLATY-BACKED NIGHTINGALE-THRUSH (Catharus fuscater)
RUDDY-CAPPED NIGHTINGALE-THRUSH (Catharus frantzii)
BLACK-HEADED NIGHTINGALE-THRUSH (Catharus mexicanus)
SWAINSON'S THRUSH (Catharus ustulatus) [b]
WOOD THRUSH (Hylocichla mustelina) [b]
SOOTY THRUSH (Turdus nigrescens)
MOUNTAIN THRUSH (Turdus plebejus)
CLAY-COLORED THRUSH (Turdus grayi)
WHITE-THROATED THRUSH (Turdus assimilis)
Ptilogonatidae (Silky-flycatchers)
BLACK-AND-YELLOW SILKY-FLYCATCHER (Phainoptila melanoxantha)
LONG-TAILED SILKY-FLYCATCHER (Ptilogonys caudatus)
Parulidae (New World Warblers)
OVENBIRD (Seiurus aurocapilla) [b]
LOUISIANA WATERTHRUSH (Parkesia motacilla) [b]
NORTHERN WATERTHRUSH (Parkesia noveboracensis) [b]
GOLDEN-WINGED WARBLER (Vermivora chrysoptera) [b]
BLACK-AND-WHITE WARBLER (Mniotilta varia) [b]
PROTHONOTARY WARBLER (Protonotaria citrea) [b]
FLAME-THROATED WARBLER (Oreothlypis gutturalis)

Sooty-capped Chlorospingus is one of the many Chiriqui endemics we encountered in the highlands. (Photo by tour participant Chris DeCilio)
TENNESSEE WARBLER (Oreothlypis peregrina) [b] GRAY-CROWNED YELLOWTHROAT (Geothlypis poliocephala)
MOURNING WARBLER (Geothlypis philadelphia) [b]
KENTUCKY WARBLER (Geothlypis formosa) [b]
OLIVE-CROWNED YELLOWTHROAT (Geothlypis semiflava)
TROPICAL PARULA (Setophaga pitiayumi)
BLACKBURNIAN WARBLER (Setophaga fusca) [b]
YELLOW WARBLER (Setophaga petechia) [b]
YELLOW WARBLER (MANGROVE) (Setophaga petechia erithachorides)
CHESTNUT-SIDED WARBLER (Setophaga pensylvanica) [b]
TOWNSEND'S WARBLER (Setophaga townsendi) [b]
BLACK-THROATED GREEN WARBLER (Setophaga virens) [b]
RUFOUS-CAPPED WARBLER (Basileuterus rufifrons)
BLACK-CHEEKED WARBLER (Basileuterus melanogenys)
GOLDEN-CROWNED WARBLER (Basileuterus culicivorus)
THREE-STRIPED WARBLER (Basileuterus tristriatus)
BUFF-RUMPED WARBLER (Myiothlypis fulvicauda)
WILSON'S WARBLER (Cardellina pusilla) [b]
SLATE-THROATED REDSTART (Myioborus miniatus)
COLLARED REDSTART (Myioborus torquatus)
WRENTHRUSH (Zeledonia coronata)
Thraupidae (Tanagers and Allies)
GRAY-HEADED TANAGER (Eucometis penicillata)
WHITE-SHOULDERED TANAGER (Tachyphonus luctuosus)
TAWNY-CRESTED TANAGER (Tachyphonus delatrii)
WHITE-LINED TANAGER (Tachyphonus rufus)
WHITE-THROATED SHRIKE-TANAGER (Lanio leucothorax)
CRIMSON-COLLARED TANAGER (Ramphocelus sanguinolentus)
PASSERINI'S TANAGER (Ramphocelus passerinii)
CHERRIE'S TANAGER (Ramphocelus costaricensis)
BLUE-AND-GOLD TANAGER (Bangsia arcaei) [*]
BLUE-GRAY TANAGER (Thraupis episcopus)
PALM TANAGER (Thraupis palmarum)
GOLDEN-HOODED TANAGER (Tangara larvata)
SPECKLED TANAGER (Tangara guttata)
SPANGLE-CHEEKED TANAGER (Tangara dowii)
PLAIN-COLORED TANAGER (Tangara inornata)
BAY-HEADED TANAGER (Tangara gyrola)
EMERALD TANAGER (Tangara florida)
SILVER-THROATED TANAGER (Tangara icterocephala)
SCARLET-THIGHED DACNIS (Dacnis venusta)

In addition to all the great birds, this trip was also excellent for mammals. One of the best was this Northern Tamandua that joined us during our picnic breakfast at Braulio Carrillo. (Photo by tour participant Marshall Dahl)
BLUE DACNIS (Dacnis cayana) SHINING HONEYCREEPER (Cyanerpes lucidus)
RED-LEGGED HONEYCREEPER (Cyanerpes cyaneus)
GREEN HONEYCREEPER (Chlorophanes spiza)
BLACK-AND-YELLOW TANAGER (Chrysothlypis chrysomelas)
SLATY FLOWERPIERCER (Diglossa plumbea)
BLUE-BLACK GRASSQUIT (Volatinia jacarina)
VARIABLE SEEDEATER (Sporophila corvina)
WHITE-COLLARED SEEDEATER (Sporophila torqueola)
NICARAGUAN SEED-FINCH (Oryzoborus nuttingi)
THICK-BILLED SEED-FINCH (Oryzoborus funereus)
BANANAQUIT (Coereba flaveola)
YELLOW-FACED GRASSQUIT (Tiaris olivaceus)
DUSKY-FACED TANAGER (Mitrospingus cassinii)
GRAYISH SALTATOR (Saltator coerulescens)
BUFF-THROATED SALTATOR (Saltator maximus)
BLACK-HEADED SALTATOR (Saltator atriceps)
SLATE-COLORED GROSBEAK (Saltator grossus)
Emberizidae (Buntings and New World Sparrows)
YELLOW-THIGHED FINCH (Pselliophorus tibialis)
LARGE-FOOTED FINCH (Pezopetes capitalis)
SOOTY-FACED FINCH (Arremon crassirostris)
CHESTNUT-CAPPED BRUSH-FINCH (Arremon brunneinucha)
ORANGE-BILLED SPARROW (Arremon aurantiirostris)
OLIVE SPARROW (Arremonops rufivirgatus)
BLACK-STRIPED SPARROW (Arremonops conirostris)
WHITE-NAPED BRUSH-FINCH (YELLOW-THROATED) (Atlapetes albinucha gutturalis)
WHITE-EARED GROUND-SPARROW (Melozone leucotis)
PREVOST'S GROUND-SPARROW (CABANIS'S) (Melozone biarcuata cabanisi)
STRIPE-HEADED SPARROW (Peucaea ruficauda)
RUFOUS-COLLARED SPARROW (Zonotrichia capensis)
VOLCANO JUNCO (Junco vulcani)
COMMON CHLOROSPINGUS (Chlorospingus flavopectus)
SOOTY-CAPPED CHLOROSPINGUS (Chlorospingus pileatus)
Cardinalidae (Cardinals and Allies)
HEPATIC TANAGER (Piranga flava)
SUMMER TANAGER (Piranga rubra) [b]
FLAME-COLORED TANAGER (Piranga bidentata)
WHITE-WINGED TANAGER (Piranga leucoptera)
RED-THROATED ANT-TANAGER (Habia fuscicauda)
CARMIOL'S TANAGER (Chlorothraupis carmioli)

Headlining an impressive list of herps seen was this rarely seen, canopy-dwelling Neotropical Chameleon, which we found dozing next to the suspension bridge at La Selva. (Photo by guide Jay VanderGaast)
BLACK-FACED GROSBEAK (Caryothraustes poliogaster) BLACK-THIGHED GROSBEAK (Pheucticus tibialis) [*]
ROSE-BREASTED GROSBEAK (Pheucticus ludovicianus) [b]
BLUE-BLACK GROSBEAK (Cyanocompsa cyanoides)
BLUE GROSBEAK (Passerina caerulea)
Icteridae (Troupials and Allies)
RED-WINGED BLACKBIRD (Agelaius phoeniceus)
RED-BREASTED BLACKBIRD (Sturnella militaris)
EASTERN MEADOWLARK (Sturnella magna)
MELODIOUS BLACKBIRD (Dives dives)
GREAT-TAILED GRACKLE (Quiscalus mexicanus)
SHINY COWBIRD (Molothrus bonariensis)
BRONZED COWBIRD (Molothrus aeneus)
GIANT COWBIRD (Molothrus oryzivorus)
BLACK-COWLED ORIOLE (Icterus prosthemelas)
BALTIMORE ORIOLE (Icterus galbula) [b]
YELLOW-BILLED CACIQUE (Amblycercus holosericeus)
SCARLET-RUMPED CACIQUE (SCARLET-RUMPED) (Cacicus uropygialis microrhynchus)
CHESTNUT-HEADED OROPENDOLA (Psarocolius wagleri)
MONTEZUMA OROPENDOLA (Psarocolius montezuma)
Fringillidae (Siskins, Crossbills, and Allies)
SCRUB EUPHONIA (Euphonia affinis)
YELLOW-CROWNED EUPHONIA (Euphonia luteicapilla)
YELLOW-THROATED EUPHONIA (Euphonia hirundinacea)
ELEGANT EUPHONIA (Euphonia elegantissima)
SPOT-CROWNED EUPHONIA (Euphonia imitans)
OLIVE-BACKED EUPHONIA (Euphonia gouldi)
TAWNY-CAPPED EUPHONIA (Euphonia anneae)
GOLDEN-BROWED CHLOROPHONIA (Chlorophonia callophrys)
YELLOW-BELLIED SISKIN (Spinus xanthogastrus)
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus) [I]
LONG-NOSED BAT (Rhynchonycteris naso)
NORTHERN GHOST BAT (Diclidurus albus)
MANTLED HOWLER MONKEY (Alouatta palliata)
WHITE-THROATED CAPUCHIN (Cebus capucinus)
CENTRAL AMERICAN SPIDER MONKEY (Ateles geoffroyi)
HOFFMANN'S TWO-TOED SLOTH (Choloepus hoffmanni)
NORTHERN TAMANDUA (Tamandua mexicana)
VARIEGATED SQUIRREL (Sciurus variegatoides)
RED-TAILED SQUIRREL (Sciurus granatensis)
ALFARO'S PYGMY SQUIRREL (Microsciurus alfari)
SUMICHRAST'S VESPER RAT (Nyctomys sumichrasti)
MEXICAN HAIRY PORCUPINE (Coendou mexicanus)
CENTRAL AMERICAN AGOUTI (Dasyprocta punctata)
NORTHERN RACCOON (Procyon lotor)
WHITE-NOSED COATI (Nasua narica)
LONG-TAILED WEASEL (Mustela frenata)
TAYRA (Eira barbara)
COLLARED PECCARY (Tayassu tajacu)
WHITE-TAILED DEER (Odocoileus virginianus)
Herps
SLENDER ANOLE (Anolis limifrons)
LEMUR ANOLE (Norops lemurinus)
STREAM ANOLE (Anolis poecilopus)
PUG-NOSED ANOLE (Norops capito)
GREEN TREE ANOLE (Norops biporcatus)
NEOTROPICAL CHAMELEON (Polychrus gutturosus)
GREEN IGUANA (Iguana iguana)
BLACK SPINY-TAILED IGUANA (Ctenosaura similis)
COMMON BASILISK (Basiliscus basiliscus)
GREEN BASILISK (Basiliscus plumifrons)
STRIPED BASILISK (Basiliscus vittatus)
TURNIP-TAILED GECKO (Thecadactylus rapicauda)
TROPICAL HOUSE GECKO (Hemidactylus mabouia)
CENTRAL AMERICAN WHIPTAIL (Ameiva festiva)
YELLOW-SPOTTED NIGHT LIZARD (Lepidophyma flavimaculatum)
FALSE FER-DE-LANCE (Xenodon rabdocephalus)
GREEN PARROT SNAKE (Leptophis ahaetulla)
BROWN BLUNT-HEADED VINE SNAKE (Imantodes cenchoa)
AMERICAN CROCODILE (Crocodylus acutus)
SPECTACLED CAIMAN (Caiman crocodilus)
CANE TOAD (Bufo marinus)
SMOOTH-SKINNED TOAD (Bufo haematicus)
SMOKY JUNGLE FROG (Leptodactylus pentadactylus)
STRAWBERRY POISON DART FROG (Dendrobates pumilio)
MASKED TREE FROG (Smilisca phaeota)
MEXICAN TREE FROG (Smilisca baudinii)
BLACK RIVER TURTLE (Rhinoclemmys funereal)
Totals for the tour: 502 bird taxa and 19 mammal taxa