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This superb male Black-crested Coquette was a siesta-time find at the verbena hedges outside our lodge at Arenal. (Photo by Joe Suchecki)
When I was first asked to put this tour together, I was pretty excited, as I had some great memories from my last Kania trip in eastern Venezuela a few years back. But I was a little concerned that, given the late date, and it being Costa Rica's high season, we might not be able to pull it all together in a way that was suitable. Well, I was very pleased that, with the help of our excellent ground agents in Costa Rica, we were able to get this thing off the ground and running. It turned out to be a very good trip, too, with lots of ground covered, and a whole lot of super birds along the way!
We started off in the highlands of the Savegre Valley where the conditions were warm and dry, and the birding was hot! Though our quetzal experience wasn't the best one I've ever enjoyed, a number of other specialties more than made up for that. Among the stellar performers here were: a trio of Spotted Wood-Quail feeding quietly in the oak forest almost too close for binoculars; the boldest Buffy-crowned Wood-Partridge I have ever seen, singing out in the open after several furtive glances at the group; a sneaky Bare-shanked Screech-Owl that almost, but not quite, gave us the slip; a cooperative pair of Buffy Tuftedcheeks along the Providencia Road; the sleek and elegant Long-tailed Silky-Flycatchers appeasing Denis's craving for this species; and a fantastically curious Wrenthrush that paraded out in the open for all to see to their satisfaction!
From the highlands we moved south to San Isidro, and Los Cusingos reserve, where we enjoyed day roosting Mottled Owl and Common Potoo, a small army ant swarm with Bicolored Antbirds, Tawny-winged Woodcreepers, and Gray-headed Tanagers, and a moonwalking Red-capped Manakin among many other species. Further south still, the area around Wilson Botanical Gardens gave us a bunch of southern specialties, from a stunning male White-crested Coquette right outside our cabins, to Snowy-bellied Hummingbird, Masked Yellowthroat, and an excited pair of Costa Rican Brush-Finches.
The south Pacific lowlands were up next, where another bunch of specialties awaited us. The key bird here was the endemic Black-cheeked Ant-Tanager, which we saw beautifully along the Golfito road. Other highlights in this region included a perched Tiny Hawk (great spotting Jon!), a Gray-lined Hawk with the tail end of an iguana in its talons, several Veraguan Mangos and a Sapphire-throated Hummingbird, both recent arrivals in the country from Panama, our only pair of Fiery-billed Aracaris for the trip (where were they all?), lovely Golden-naped Woodpeckers, the striking Black-striped Woodcreeper, and great looks at both Turquoise and Yellow-billed cotingas.
Up the coast, we made a brief stop in the Carara region and a morning foray into the dry Guanacaste lowlands, where we tallied northwestern specialties like Double-striped Thick-knee, Plain-capped Starthroat, Cinnamon Hummingbird, the exquisite Turquoise-browed Motmot, Orange-fronted Parakeet, several flycatchers, White-throated Magpie-Jay, and White-lored Gnatcatcher, though the Lesser Ground-Cuckoos stayed frustratingly out of sight. Then it was onward to the northern half of the country, with the now quiet Arenal Volcano as our first stop.
Arenal was alive with birds, and though there aren't too many regional specialties, there are a lot of great birds to be found. Antbirds were a highlight, particularly at our big ant swarm at the Sky Walk, where we had multiple Spotted, Bicolored, Zeledon's, and the superb Ocellated antbirds all at point-blank range for some awesome views. Easily one of the best ant swarms I've seen in the country! We also had wonderful views of Dull-mantled and Bare-crowned antbirds, and an exceptionally easy Thicket Antpitta, for one of the best antbird days imaginable. Other birds that made our time here memorable included Great Curassow, Black-crested Coquette, a pair of White-fronted Nunbirds, great views of a very local Keel-billed Motmot, a distant, but glowing, Lovely Cotinga, Black-headed Nightingale-Thrush, Song Wren, and many more.
We wrapped things up with quick visits to three locales. Our boat ride at Cano Negro was superb, despite the low water levels, and views of a close male Sungrebe, an even closer American Pygmy Kingfisher, several of the extremely local Nicaraguan Grackles, and a host of other wetland birds made it a memorable one. Our night drive there was productive too, with a couple of striking Striped Owls, a pair of Pacific Screech-Owls, and a Black-and-white Owl picking off a bat under a streetlight making for an exciting night indeed! Bosque de Paz was also nice, with a couple of Scaled Antpittas hopping down the trail ahead of us, a Great Black-Hawk flying out of its nest tree, and, at last, some inexplicably scarce but gorgeous Spangle-cheeked Tanagers. And finally, we finished with a buzz of hummers at La Paz, including the tiny endemic Coppery-headed Emerald and hefty Violet Sabrewings.
All in all, this trip was a lot of fun, and I really enjoyed the camaraderie, good humor, and enthusiasm you all showed throughout the trip. I'd be quite happy if all my groups came with the group dynamics already well established! I hope spring has finally found you all, as it seems to have finally done here. And I hope another Kania trip comes my way someday soon. I'd look forward to seeing any or all of you on another trip in the near future.
Cheers,
--Jay
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Tinamidae (Tinamous)

With the recent split of Gray Hawk, we had an extra species to look for, as Costa Rica lies right in the contact zone between the two species. We saw both: a Gray-lined Hawk in the Coto 47 Marsh area, and this Gray Hawk along the roadside near Cano Negro. (Photo by guide Jay VanderGaast)
GREAT TINAMOU (Tinamus major) LITTLE TINAMOU (Crypturellus soui) [*]
Anatidae (Ducks, Geese, and Waterfowl)
BLACK-BELLIED WHISTLING-DUCK (Dendrocygna autumnalis)
MUSCOVY DUCK (Cairina moschata)
AMERICAN WIGEON (Anas americana) [b]
BLUE-WINGED TEAL (Anas discors) [b]
Cracidae (Guans, Chachalacas, and Curassows)
GRAY-HEADED CHACHALACA (Ortalis cinereiceps)
CRESTED GUAN (Penelope purpurascens)
BLACK GUAN (Chamaepetes unicolor)
GREAT CURASSOW (Crax rubra)
Odontophoridae (New World Quail)
BUFFY-CROWNED WOOD-PARTRIDGE (Dendrortyx leucophrys)
SPOTTED WOOD-QUAIL (Odontophorus guttatus)
Podicipedidae (Grebes)
LEAST GREBE (Tachybaptus dominicus)
Ciconiidae (Storks)
WOOD STORK (Mycteria americana)
Fregatidae (Frigatebirds)
MAGNIFICENT FRIGATEBIRD (Fregata magnificens)
Phalacrocoracidae (Cormorants and Shags)
NEOTROPIC CORMORANT (Phalacrocorax brasilianus)
Anhingidae (Anhingas)
ANHINGA (Anhinga anhinga)
Pelecanidae (Pelicans)
BROWN PELICAN (Pelecanus occidentalis)
Ardeidae (Herons, Egrets, and Bitterns)
PINNATED BITTERN (Botaurus pinnatus)
FASCIATED TIGER-HERON (Tigrisoma fasciatum)
BARE-THROATED TIGER-HERON (Tigrisoma mexicanum)
GREAT BLUE HERON (Ardea herodias)
GREAT EGRET (Ardea alba)
SNOWY EGRET (Egretta thula)
LITTLE BLUE HERON (Egretta caerulea)
TRICOLORED HERON (Egretta tricolor)
CATTLE EGRET (Bubulcus ibis)
GREEN HERON (Butorides virescens)
BLACK-CROWNED NIGHT-HERON (Nycticorax nycticorax)
BOAT-BILLED HERON (Cochlearius cochlearius)
Threskiornithidae (Ibises and Spoonbills)
WHITE IBIS (Eudocimus albus)
GLOSSY IBIS (Plegadis falcinellus)
GREEN IBIS (Mesembrinibis cayennensis)
ROSEATE SPOONBILL (Platalea ajaja)
Cathartidae (New World Vultures)

While it may no longer be actively erupting, Arenal Volcano is still a beautiful sight, and the birding on its lower slopes is pretty awesome, too! (Photo by guide Jay VanderGaast)
BLACK VULTURE (Coragyps atratus) TURKEY VULTURE (Cathartes aura)
LESSER YELLOW-HEADED VULTURE (Cathartes burrovianus)
KING VULTURE (Sarcoramphus papa)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus)
Accipitridae (Hawks, Eagles, and Kites)
PEARL KITE (Gampsonyx swainsonii)
WHITE-TAILED KITE (Elanus leucurus)
GRAY-HEADED KITE (Leptodon cayanensis)
SWALLOW-TAILED KITE (Elanoides forficatus)
DOUBLE-TOOTHED KITE (Harpagus bidentatus)
PLUMBEOUS KITE (Ictinia plumbea) [a]
TINY HAWK (Accipiter superciliosus)
CRANE HAWK (Geranospiza caerulescens)
COMMON BLACK-HAWK (MANGROVE) (Buteogallus anthracinus subtilis)
GREAT BLACK-HAWK (Buteogallus urubitinga) [N]
ROADSIDE HAWK (Rupornis magnirostris)
WHITE HAWK (Pseudastur albicollis)
SEMIPLUMBEOUS HAWK (Leucopternis semiplumbeus)
BROAD-WINGED HAWK (Buteo platypterus) [b]
GRAY HAWK (Buteo plagiatus)
GRAY-LINED HAWK (Buteo nitidus)
SHORT-TAILED HAWK (Buteo brachyurus)
SWAINSON'S HAWK (Buteo swainsoni) [b]
ZONE-TAILED HAWK (Buteo albonotatus)
RED-TAILED HAWK (COSTA RICAN) (Buteo jamaicensis costaricensis)
Rallidae (Rails, Gallinules, and Coots)
WHITE-THROATED CRAKE (Laterallus albigularis)
GRAY-BREASTED CRAKE (Laterallus exilis) [*]
GRAY-NECKED WOOD-RAIL (Aramides cajaneus)
PURPLE GALLINULE (Porphyrio martinicus)
COMMON GALLINULE (Gallinula galeata)
Heliornithidae (Finfoots)
SUNGREBE (Heliornis fulica)
Aramidae (Limpkin)
LIMPKIN (Aramus guarauna)
Burhinidae (Thick-knees)
DOUBLE-STRIPED THICK-KNEE (Burhinus bistriatus)
Recurvirostridae (Stilts and Avocets)
BLACK-NECKED STILT (Himantopus mexicanus)
Charadriidae (Plovers and Lapwings)
SOUTHERN LAPWING (Vanellus chilensis)
SEMIPALMATED PLOVER (Charadrius semipalmatus) [b]
KILLDEER (Charadrius vociferus) [b]
Jacanidae (Jacanas)
NORTHERN JACANA (Jacana spinosa)
Scolopacidae (Sandpipers and Allies)
SPOTTED SANDPIPER (Actitis macularius) [b]
SOLITARY SANDPIPER (Tringa solitaria) [b]
GREATER YELLOWLEGS (Tringa melanoleuca) [b]
WILLET (Tringa semipalmata) [b]
LESSER YELLOWLEGS (Tringa flavipes) [b]

One of 40 species of hummingbirds on the trip, the Green Violetear is among the most vocal: they could be heard chirping throughout the day in the Savegre Valley. (Photo by Joe Suchecki)
WHIMBREL (Numenius phaeopus) [b] STILT SANDPIPER (Calidris himantopus) [b]
LEAST SANDPIPER (Calidris minutilla) [b]
SEMIPALMATED SANDPIPER (Calidris pusilla) [b]
WILSON'S SNIPE (Gallinago delicata) [b]
Laridae (Gulls, Terns, and Skimmers)
LAUGHING GULL (Leucophaeus atricilla)
ROYAL TERN (Thalasseus maximus)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
PALE-VENTED PIGEON (Patagioenas cayennensis)
SCALED PIGEON (Patagioenas speciosa)
RED-BILLED PIGEON (Patagioenas flavirostris)
BAND-TAILED PIGEON (Patagioenas fasciata)
RUDDY PIGEON (Patagioenas subvinacea)
SHORT-BILLED PIGEON (Patagioenas nigrirostris)
WHITE-WINGED DOVE (Zenaida asiatica)
INCA DOVE (Columbina inca)
COMMON GROUND-DOVE (Columbina passerina)
PLAIN-BREASTED GROUND-DOVE (Columbina minuta)
RUDDY GROUND-DOVE (Columbina talpacoti)
BLUE GROUND-DOVE (Claravis pretiosa)
WHITE-TIPPED DOVE (Leptotila verreauxi)
GRAY-CHESTED DOVE (Leptotila cassini)
Cuculidae (Cuckoos)
SQUIRREL CUCKOO (Piaya cayana)
MANGROVE CUCKOO (Coccyzus minor)
STRIPED CUCKOO (Tapera naevia)
LESSER GROUND-CUCKOO (Morococcyx erythropygus) [*]
SMOOTH-BILLED ANI (Crotophaga ani)
GROOVE-BILLED ANI (Crotophaga sulcirostris)
Strigidae (Owls)
PACIFIC SCREECH-OWL (Megascops cooperi)
TROPICAL SCREECH-OWL (Megascops choliba)
BARE-SHANKED SCREECH-OWL (Megascops clarkii)

One of the many highlights of our boat ride at Cano Negro was this very confiding male American Pygmy Kingfisher. (Photo by Joe Suchecki.)
SPECTACLED OWL (Pulsatrix perspicillata) COSTA RICAN PYGMY-OWL (Glaucidium costaricanum)
FERRUGINOUS PYGMY-OWL (Glaucidium brasilianum)
MOTTLED OWL (Ciccaba virgata)
BLACK-AND-WHITE OWL (Ciccaba nigrolineata)
STRIPED OWL (Pseudoscops clamator)
UNSPOTTED SAW-WHET OWL (Aegolius ridgwayi) [*]
Caprimulgidae (Nightjars and Allies)
LESSER NIGHTHAWK (Chordeiles acutipennis)
COMMON PAURAQUE (Nyctidromus albicollis)
DUSKY NIGHTJAR (Antrostomus saturatus)
Nyctibiidae (Potoos)
COMMON POTOO (Nyctibius griseus)
Apodidae (Swifts)
CHESTNUT-COLLARED SWIFT (Streptoprocne rutila)
WHITE-COLLARED SWIFT (Streptoprocne zonaris)
VAUX'S SWIFT (Chaetura vauxi)
COSTA RICAN SWIFT (Chaetura fumosa)
Trochilidae (Hummingbirds)
WHITE-NECKED JACOBIN (Florisuga mellivora)
BAND-TAILED BARBTHROAT (Threnetes ruckeri)
GREEN HERMIT (Phaethornis guy)
LONG-BILLED HERMIT (Phaethornis longirostris)
STRIPE-THROATED HERMIT (Phaethornis striigularis)
BROWN VIOLETEAR (Colibri delphinae)
GREEN VIOLETEAR (Colibri thalassinus)
PURPLE-CROWNED FAIRY (Heliothryx barroti)

We were a bit confused by the crest color of this male White-crested Coquette, until we realized that what we thought was the crest was actually the projections from the side of its dark gorget! (Photo by Joe Suchecki)
GREEN-BREASTED MANGO (Anthracothorax prevostii) VERAGUAN MANGO (Anthracothorax veraguensis)
GREEN THORNTAIL (Discosura conversii)
BLACK-CRESTED COQUETTE (Lophornis helenae)
WHITE-CRESTED COQUETTE (Lophornis adorabilis)
GREEN-CROWNED BRILLIANT (Heliodoxa jacula)
MAGNIFICENT HUMMINGBIRD (Eugenes fulgens)
LONG-BILLED STARTHROAT (Heliomaster longirostris)
PLAIN-CAPPED STARTHROAT (Heliomaster constantii)
FIERY-THROATED HUMMINGBIRD (Panterpe insignis)
PURPLE-THROATED MOUNTAIN-GEM (Lampornis calolaemus)
WHITE-THROATED MOUNTAIN-GEM (GRAY-TAILED) (Lampornis castaneoventris cinereicauda)
MAGENTA-THROATED WOODSTAR (Calliphlox bryantae)
RUBY-THROATED HUMMINGBIRD (Archilochus colubris) [b]
VOLCANO HUMMINGBIRD (Selasphorus flammula)
SCINTILLANT HUMMINGBIRD (Selasphorus scintilla)
GARDEN EMERALD (Chlorostilbon assimilis)
VIOLET-HEADED HUMMINGBIRD (Klais guimeti)
SCALY-BREASTED HUMMINGBIRD (Phaeochroa cuvierii)
VIOLET SABREWING (Campylopterus hemileucurus)
BRONZE-TAILED PLUMELETEER (Chalybura urochrysia) [*]
CROWNED WOODNYMPH (Thalurania colombica)
STRIPE-TAILED HUMMINGBIRD (Eupherusa eximia)
BLACK-BELLIED HUMMINGBIRD (Eupherusa nigriventris)
WHITE-TAILED EMERALD (Elvira chionura)
COPPERY-HEADED EMERALD (Elvira cupreiceps) [E]
CHARMING HUMMINGBIRD (Amazilia decora)
STEELY-VENTED HUMMINGBIRD (Amazilia saucerrottei)
SNOWY-BELLIED HUMMINGBIRD (Amazilia edward)
RUFOUS-TAILED HUMMINGBIRD (Amazilia tzacatl)
CINNAMON HUMMINGBIRD (Amazilia rutila)
SAPPHIRE-THROATED HUMMINGBIRD (Lepidopyga coeruleogularis)
BLUE-THROATED GOLDENTAIL (Hylocharis eliciae)
Trogonidae (Trogons)
RESPLENDENT QUETZAL (Pharomachrus mocinno)

The gang in the gorgeous montane oak forest above Savegre Lodge. (Photo by guide Jay VanderGaast)
SLATY-TAILED TROGON (Trogon massena) BLACK-HEADED TROGON (Trogon melanocephalus)
BAIRD'S TROGON (Trogon bairdii)
GARTERED TROGON (Trogon caligatus)
BLACK-THROATED TROGON (Trogon rufus)
ORANGE-BELLIED TROGON (Trogon aurantiiventris)
COLLARED TROGON (Trogon collaris)
Momotidae (Motmots)
BLUE-CROWNED MOTMOT (LESSON'S) (Momotus coeruliceps lessonii)
KEEL-BILLED MOTMOT (Electron carinatum)
BROAD-BILLED MOTMOT (Electron platyrhynchum)
TURQUOISE-BROWED MOTMOT (Eumomota superciliosa)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)
AMAZON KINGFISHER (Chloroceryle amazona)
GREEN KINGFISHER (Chloroceryle americana)
AMERICAN PYGMY KINGFISHER (Chloroceryle aenea)
Bucconidae (Puffbirds)
WHITE-NECKED PUFFBIRD (Notharchus hyperrhynchus)
PIED PUFFBIRD (Notharchus tectus)
WHITE-FRONTED NUNBIRD (Monasa morphoeus)
Galbulidae (Jacamars)
RUFOUS-TAILED JACAMAR (Galbula ruficauda)
Semnornithidae (Toucan-Barbets)
PRONG-BILLED BARBET (Semnornis frantzii)
Ramphastidae (Toucans)
COLLARED ARACARI (Pteroglossus torquatus)
FIERY-BILLED ARACARI (Pteroglossus frantzii)
BLACK-MANDIBLED TOUCAN (CHESTNUT-MANDIBLED) (Ramphastos ambiguus swainsonii)
KEEL-BILLED TOUCAN (Ramphastos sulfuratus)
Picidae (Woodpeckers)
OLIVACEOUS PICULET (Picumnus olivaceus)
ACORN WOODPECKER (Melanerpes formicivorus)
GOLDEN-NAPED WOODPECKER (Melanerpes chrysauchen)
BLACK-CHEEKED WOODPECKER (Melanerpes pucherani)
RED-CROWNED WOODPECKER (Melanerpes rubricapillus)

What may have been my best army ant swarm ever at the Arenal Sky Walk reserve featured close views of 4 species of antbirds, including several amazing Ocellated Antbirds, the Holy Grail of all CR antbirds! (Photo by guide Jay VanderGaast)
HOFFMANN'S WOODPECKER (Melanerpes hoffmannii) SMOKY-BROWN WOODPECKER (Picoides fumigatus)
HAIRY WOODPECKER (Picoides villosus)
RUFOUS-WINGED WOODPECKER (Piculus simplex)
GOLDEN-OLIVE WOODPECKER (Colaptes rubiginosus)
LINEATED WOODPECKER (Dryocopus lineatus)
PALE-BILLED WOODPECKER (Campephilus guatemalensis)
Falconidae (Falcons and Caracaras)
BARRED FOREST-FALCON (Micrastur ruficollis) [*]
CRESTED CARACARA (Caracara cheriway)
YELLOW-HEADED CARACARA (Milvago chimachima)
LAUGHING FALCON (Herpetotheres cachinnans)
AMERICAN KESTREL (Falco sparverius) [b]
PEREGRINE FALCON (Falco peregrinus) [b]
Psittacidae (Parrots)
SULPHUR-WINGED PARAKEET (Pyrrhura hoffmanni)
CRIMSON-FRONTED PARAKEET (Aratinga finschi)
OLIVE-THROATED PARAKEET (Aratinga nana)
ORANGE-FRONTED PARAKEET (Aratinga canicularis)
BROWN-THROATED PARAKEET (Aratinga pertinax)
SCARLET MACAW (Ara macao)
BARRED PARAKEET (Bolborhynchus lineola)
ORANGE-CHINNED PARAKEET (Brotogeris jugularis)
BROWN-HOODED PARROT (Pyrilia haematotis)
BLUE-HEADED PARROT (Pionus menstruus)
WHITE-CROWNED PARROT (Pionus senilis)
RED-LORED PARROT (Amazona autumnalis)
YELLOW-NAPED PARROT (Amazona auropalliata) [*]
WHITE-FRONTED PARROT (Amazona albifrons)
MEALY PARROT (Amazona farinosa)
Thamnophilidae (Typical Antbirds)
GREAT ANTSHRIKE (Taraba major)
BARRED ANTSHRIKE (Thamnophilus doliatus)

Another member of the ant swarm attendees, the dapper little Spotted Antbird. (Photo by Joe Suchecki)
BLACK-HOODED ANTSHRIKE (Thamnophilus bridgesi) RUSSET ANTSHRIKE (Thamnistes anabatinus)
PLAIN ANTVIREO (Dysithamnus mentalis)
STREAK-CROWNED ANTVIREO (Dysithamnus striaticeps)
SLATY ANTWREN (Myrmotherula schisticolor)
DOT-WINGED ANTWREN (Microrhopias quixensis)
DUSKY ANTBIRD (Cercomacra tyrannina)
BARE-CROWNED ANTBIRD (Gymnocichla nudiceps)
CHESTNUT-BACKED ANTBIRD (Myrmeciza exsul)
DULL-MANTLED ANTBIRD (Myrmeciza laemosticta)
ZELEDON'S ANTBIRD (Myrmeciza zeledoni)
BICOLORED ANTBIRD (Gymnopithys leucaspis bicolor)
SPOTTED ANTBIRD (Hylophylax naevioides)
OCELLATED ANTBIRD (Phaenostictus mcleannani)
Grallariidae (Antpittas)
SCALED ANTPITTA (Grallaria guatimalensis)
THICKET ANTPITTA (Hylopezus dives)
Rhinocryptidae (Tapaculos)
SILVERY-FRONTED TAPACULO (Scytalopus argentifrons)
Formicariidae (Antthrushes)
BLACK-FACED ANTTHRUSH (Formicarius analis)
Furnariidae (Ovenbirds and Woodcreepers)
OLIVACEOUS WOODCREEPER (Sittasomus griseicapillus)
TAWNY-WINGED WOODCREEPER (Dendrocincla anabatina)
WEDGE-BILLED WOODCREEPER (Glyphorynchus spirurus)
NORTHERN BARRED-WOODCREEPER (Dendrocolaptes sanctithomae)
COCOA WOODCREEPER (Xiphorhynchus susurrans)
BLACK-STRIPED WOODCREEPER (Xiphorhynchus lachrymosus)
SPOTTED WOODCREEPER (Xiphorhynchus erythropygius)
BROWN-BILLED SCYTHEBILL (Campylorhamphus pusillus)
STREAK-HEADED WOODCREEPER (Lepidocolaptes souleyetii)
SPOT-CROWNED WOODCREEPER (Lepidocolaptes affinis)
PLAIN XENOPS (Xenops minutus)
BUFFY TUFTEDCHEEK (Pseudocolaptes lawrencii)
LINEATED FOLIAGE-GLEANER (Syndactyla subalaris)
STRIPED WOODHAUNTER (Hyloctistes subulatus)
BUFF-THROATED FOLIAGE-GLEANER (HYPOPHAEUS) (Automolus ochrolaemus hypophaeus)
BUFF-THROATED FOLIAGE-GLEANER (EXSERTUS) (Automolus ochrolaemus exsertus)
RUDDY FOLIAGE-GLEANER (Automolus rubiginosus)
SPOTTED BARBTAIL (Premnoplex brunnescens)
RUDDY TREERUNNER (Margarornis rubiginosus)

A common bird of the highlands, the Sooty Thrush is one of a number of Chiriqui endemics, restricted to the mountains of Costa Rica and Panama. (Photo by Joe Suchecki)
RED-FACED SPINETAIL (Cranioleuca erythrops) PALE-BREASTED SPINETAIL (Synallaxis albescens)
SLATY SPINETAIL (Synallaxis brachyura)
Tyrannidae (Tyrant Flycatchers)
NORTHERN BEARDLESS-TYRANNULET (Camptostoma imberbe)
SOUTHERN BEARDLESS-TYRANNULET (Camptostoma obsoletum)
YELLOW TYRANNULET (Capsiempis flaveola)
YELLOW-CROWNED TYRANNULET (Tyrannulus elatus)
GREENISH ELAENIA (Myiopagis viridicata)
YELLOW-BELLIED ELAENIA (Elaenia flavogaster)
LESSER ELAENIA (Elaenia chiriquensis)
MOUNTAIN ELAENIA (Elaenia frantzii)
TORRENT TYRANNULET (Serpophaga cinerea)
OCHRE-BELLIED FLYCATCHER (Mionectes oleagineus)
SLATY-CAPPED FLYCATCHER (Leptopogon superciliaris)
PALTRY TYRANNULET (Zimmerius vilissimus)
SCALE-CRESTED PYGMY-TYRANT (Lophotriccus pileatus)
NORTHERN BENTBILL (Oncostoma cinereigulare) [*]
SLATE-HEADED TODY-FLYCATCHER (Poecilotriccus sylvia) [*]
COMMON TODY-FLYCATCHER (Todirostrum cinereum)
BLACK-HEADED TODY-FLYCATCHER (Todirostrum nigriceps)
EYE-RINGED FLATBILL (Rhynchocyclus brevirostris)
YELLOW-OLIVE FLYCATCHER (Tolmomyias sulphurescens)
WHITE-THROATED SPADEBILL (Platyrinchus mystaceus)
GOLDEN-CROWNED SPADEBILL (Platyrinchus coronatus)
SULPHUR-RUMPED FLYCATCHER (Myiobius sulphureipygius aureatus)
TUFTED FLYCATCHER (Mitrephanes phaeocercus)
DARK PEWEE (Contopus lugubris)
EASTERN WOOD-PEWEE (Contopus virens) [b]
TROPICAL PEWEE (Contopus cinereus)
YELLOW-BELLIED FLYCATCHER (Empidonax flaviventris) [b]
WILLOW FLYCATCHER (Empidonax traillii) [b]
YELLOWISH FLYCATCHER (Empidonax flavescens)
BLACK-CAPPED FLYCATCHER (Empidonax atriceps)
BLACK PHOEBE (Sayornis nigricans)
LONG-TAILED TYRANT (Colonia colonus)
BRIGHT-RUMPED ATTILA (Attila spadiceus)
RUFOUS MOURNER (Rhytipterna holerythra)
DUSKY-CAPPED FLYCATCHER (Myiarchus tuberculifer)
NUTTING'S FLYCATCHER (Myiarchus nuttingi)
GREAT CRESTED FLYCATCHER (Myiarchus crinitus) [b]
BROWN-CRESTED FLYCATCHER (Myiarchus tyrannulus)
GREAT KISKADEE (Pitangus sulphuratus)
BOAT-BILLED FLYCATCHER (Megarynchus pitangua)
SOCIAL FLYCATCHER (Myiozetetes similis)
GRAY-CAPPED FLYCATCHER (Myiozetetes granadensis)
WHITE-RINGED FLYCATCHER (Conopias albovittatus)
GOLDEN-BELLIED FLYCATCHER (Myiodynastes hemichrysus)
STREAKED FLYCATCHER (Myiodynastes maculatus)
PIRATIC FLYCATCHER (Legatus leucophaius) [a]
TROPICAL KINGBIRD (Tyrannus melancholicus)
SCISSOR-TAILED FLYCATCHER (Tyrannus forficatus) [b]
FORK-TAILED FLYCATCHER (Tyrannus savana)
Cotingidae (Cotingas)
LOVELY COTINGA (Cotinga amabilis)
TURQUOISE COTINGA (Cotinga ridgwayi)
RUFOUS PIHA (Lipaugus unirufus)
YELLOW-BILLED COTINGA (Carpodectes antoniae)
SNOWY COTINGA (Carpodectes nitidus)
Pipridae (Manakins)
WHITE-RUFFED MANAKIN (Corapipo altera)

This one's for you, Denis! Not the best photo, but still a stellar bird: Long-tailed Silky-Flycatcher. (Photo by guide Jay VanderGaast)
LONG-TAILED MANAKIN (Chiroxiphia linearis) RED-CAPPED MANAKIN (Ceratopipra mentalis)
WHITE-COLLARED MANAKIN (Manacus candei)
ORANGE-COLLARED MANAKIN (Manacus aurantiacus)
BLUE-CROWNED MANAKIN (Lepidothrix coronata)
Tityridae (Tityras and Allies)
BLACK-CROWNED TITYRA (Tityra inquisitor)
MASKED TITYRA (Tityra semifasciata)
BARRED BECARD (Pachyramphus versicolor)
CINNAMON BECARD (Pachyramphus cinnamomeus)
ROSE-THROATED BECARD (Pachyramphus aglaiae)
Vireonidae (Vireos)
YELLOW-THROATED VIREO (Vireo flavifrons) [b]
BLUE-HEADED VIREO (Vireo solitarius) [b]
YELLOW-WINGED VIREO (Vireo carmioli)
BROWN-CAPPED VIREO (Vireo leucophrys)
PHILADELPHIA VIREO (Vireo philadelphicus) [b]
TAWNY-CROWNED GREENLET (Hylophilus ochraceiceps)
LESSER GREENLET (Hylophilus decurtatus)
GREEN SHRIKE-VIREO (Vireolanius pulchellus)
RUFOUS-BROWED PEPPERSHRIKE (Cyclarhis gujanensis)
Corvidae (Crows, Jays, and Magpies)
AZURE-HOODED JAY (Cyanolyca cucullata) [*]
WHITE-THROATED MAGPIE-JAY (Calocitta formosa)
BROWN JAY (Psilorhinus morio)
Hirundinidae (Swallows)
BLUE-AND-WHITE SWALLOW (Pygochelidon cyanoleuca)
NORTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx serripennis)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)
GRAY-BREASTED MARTIN (Progne chalybea)
MANGROVE SWALLOW (Tachycineta albilinea)
BARN SWALLOW (Hirundo rustica) [b]
Troglodytidae (Wrens)
NIGHTINGALE WREN (Microcerculus philomela)
SCALY-BREASTED WREN (WHISTLING) (Microcerculus marginatus luscinia) [*]
HOUSE WREN (Troglodytes aedon)
OCHRACEOUS WREN (Troglodytes ochraceus)
TIMBERLINE WREN (Thryorchilus browni)
SEDGE WREN (Cistothorus platensis)
BAND-BACKED WREN (Campylorhynchus zonatus)
RUFOUS-NAPED WREN (Campylorhynchus rufinucha)
BLACK-BELLIED WREN (Pheugopedius fasciatoventris)
RUFOUS-BREASTED WREN (Pheugopedius rutilus)
SPOT-BREASTED WREN (Pheugopedius maculipectus)
BLACK-THROATED WREN (Pheugopedius atrogularis)
BANDED WREN (Thryophilus pleurostictus)
STRIPE-BREASTED WREN (Cantorchilus thoracicus)
PLAIN WREN (Cantorchilus modestus)
RIVERSIDE WREN (Cantorchilus semibadius)
BAY WREN (Cantorchilus nigricapillus)
WHITE-BREASTED WOOD-WREN (Henicorhina leucosticta)
GRAY-BREASTED WOOD-WREN (Henicorhina leucophrys)
SONG WREN (Cyphorhinus phaeocephalus)
Polioptilidae (Gnatcatchers)
TAWNY-FACED GNATWREN (Microbates cinereiventris)
LONG-BILLED GNATWREN (Ramphocaenus melanurus)
WHITE-LORED GNATCATCHER (Polioptila albiloris)
TROPICAL GNATCATCHER (Polioptila plumbea)
Cinclidae (Dippers)
AMERICAN DIPPER (Cinclus mexicanus)
Turdidae (Thrushes and Allies)
BLACK-FACED SOLITAIRE (Myadestes melanops)
BLACK-BILLED NIGHTINGALE-THRUSH (Catharus gracilirostris)

The charismatic and photogenic "Amigo de hombre", better known as the Collared Redstart. This friendly little guy posed for us on the grounds of Trogon Lodge. (Photo by guide Jay VanderGaast)
ORANGE-BILLED NIGHTINGALE-THRUSH (Catharus aurantiirostris) SLATY-BACKED NIGHTINGALE-THRUSH (Catharus fuscater)
RUDDY-CAPPED NIGHTINGALE-THRUSH (Catharus frantzii)
BLACK-HEADED NIGHTINGALE-THRUSH (Catharus mexicanus)
SWAINSON'S THRUSH (Catharus ustulatus) [b]
SOOTY THRUSH (Turdus nigrescens)
MOUNTAIN THRUSH (Turdus plebejus)
CLAY-COLORED THRUSH (Turdus grayi)
WHITE-THROATED THRUSH (Turdus assimilis)
Mimidae (Mockingbirds and Thrashers)
TROPICAL MOCKINGBIRD (Mimus gilvus)
Ptilogonatidae (Silky-flycatchers)
BLACK-AND-YELLOW SILKY-FLYCATCHER (Phainoptila melanoxantha)
LONG-TAILED SILKY-FLYCATCHER (Ptilogonys caudatus)
Parulidae (New World Warblers)
LOUISIANA WATERTHRUSH (Parkesia motacilla) [b]
NORTHERN WATERTHRUSH (Parkesia noveboracensis) [b]
GOLDEN-WINGED WARBLER (Vermivora chrysoptera) [b]
BLACK-AND-WHITE WARBLER (Mniotilta varia) [b]
PROTHONOTARY WARBLER (Protonotaria citrea) [b]
FLAME-THROATED WARBLER (Oreothlypis gutturalis)
TENNESSEE WARBLER (Oreothlypis peregrina) [b]
GRAY-CROWNED YELLOWTHROAT (Geothlypis poliocephala)
MASKED YELLOWTHROAT (CHIRIQUI) (Geothlypis aequinoctialis chiriquensis)
MOURNING WARBLER (Geothlypis philadelphia) [b]
KENTUCKY WARBLER (Geothlypis formosa) [b]
OLIVE-CROWNED YELLOWTHROAT (Geothlypis semiflava)
AMERICAN REDSTART (Setophaga ruticilla) [b]
TROPICAL PARULA (Setophaga pitiayumi)
BLACKBURNIAN WARBLER (Setophaga fusca) [b]
YELLOW WARBLER (Setophaga petechia) [b]
CHESTNUT-SIDED WARBLER (Setophaga pensylvanica) [b]
BLACK-THROATED GREEN WARBLER (Setophaga virens) [b]
RUFOUS-CAPPED WARBLER (Basileuterus rufifrons)
BLACK-CHEEKED WARBLER (Basileuterus melanogenys)
GOLDEN-CROWNED WARBLER (Basileuterus culicivorus)
THREE-STRIPED WARBLER (Basileuterus tristriatus)
BUFF-RUMPED WARBLER (Myiothlypis fulvicauda)
WILSON'S WARBLER (Cardellina pusilla) [b]
SLATE-THROATED REDSTART (Myioborus miniatus)
COLLARED REDSTART (Myioborus torquatus)
WRENTHRUSH (Zeledonia coronata)
Thraupidae (Tanagers and Allies)
GRAY-HEADED TANAGER (Eucometis penicillata)
WHITE-SHOULDERED TANAGER (Tachyphonus luctuosus)
WHITE-LINED TANAGER (Tachyphonus rufus)
WHITE-THROATED SHRIKE-TANAGER (Lanio leucothorax) [*]
CRIMSON-COLLARED TANAGER (Ramphocelus sanguinolentus) [*]
PASSERINI'S TANAGER (Ramphocelus passerinii)
CHERRIE'S TANAGER (Ramphocelus costaricensis)
BLUE-GRAY TANAGER (Thraupis episcopus)
PALM TANAGER (Thraupis palmarum)
GOLDEN-HOODED TANAGER (Tangara larvata)
SPECKLED TANAGER (Tangara guttata)
SPANGLE-CHEEKED TANAGER (Tangara dowii)
BAY-HEADED TANAGER (Tangara gyrola)
SILVER-THROATED TANAGER (Tangara icterocephala)
SCARLET-THIGHED DACNIS (Dacnis venusta)
BLUE DACNIS (Dacnis cayana)
SHINING HONEYCREEPER (Cyanerpes lucidus)
RED-LEGGED HONEYCREEPER (Cyanerpes cyaneus)
GREEN HONEYCREEPER (Chlorophanes spiza)
SLATY FLOWERPIERCER (Diglossa plumbea)
BLUE-BLACK GRASSQUIT (Volatinia jacarina)
VARIABLE SEEDEATER (Sporophila corvina)
WHITE-COLLARED SEEDEATER (Sporophila torqueola)
YELLOW-BELLIED SEEDEATER (Sporophila nigricollis)
THICK-BILLED SEED-FINCH (Oryzoborus funereus)
BANANAQUIT (Coereba flaveola)
YELLOW-FACED GRASSQUIT (Tiaris olivaceus)
STREAKED SALTATOR (Saltator striatipectus)
GRAYISH SALTATOR (Saltator coerulescens)
BUFF-THROATED SALTATOR (Saltator maximus)
BLACK-HEADED SALTATOR (Saltator atriceps)
Emberizidae (Buntings and New World Sparrows)
YELLOW-THIGHED FINCH (Pselliophorus tibialis)
LARGE-FOOTED FINCH (Pezopetes capitalis)
CHESTNUT-CAPPED BRUSH-FINCH (Arremon brunneinucha)
ORANGE-BILLED SPARROW (Arremon aurantiirostris)
COSTA RICAN BRUSH-FINCH (Arremon costaricensis)
OLIVE SPARROW (Arremonops rufivirgatus)
BLACK-STRIPED SPARROW (Arremonops conirostris)
WHITE-EARED GROUND-SPARROW (Melozone leucotis)
PREVOST'S GROUND-SPARROW (CABANIS'S) (Melozone biarcuata cabanisi)
STRIPE-HEADED SPARROW (Peucaea ruficauda)
RUFOUS-COLLARED SPARROW (Zonotrichia capensis)
VOLCANO JUNCO (Junco vulcani)
COMMON CHLOROSPINGUS (Chlorospingus flavopectus)
SOOTY-CAPPED CHLOROSPINGUS (Chlorospingus pileatus)
Cardinalidae (Cardinals and Allies)
HEPATIC TANAGER (Piranga flava)
SUMMER TANAGER (Piranga rubra) [b]
FLAME-COLORED TANAGER (Piranga bidentata)
RED-CROWNED ANT-TANAGER (Habia rubica)
RED-THROATED ANT-TANAGER (Habia fuscicauda)
BLACK-CHEEKED ANT-TANAGER (Habia atrimaxillaris) [E]
CARMIOL'S TANAGER (Chlorothraupis carmioli)
BLACK-THIGHED GROSBEAK (Pheucticus tibialis)
ROSE-BREASTED GROSBEAK (Pheucticus ludovicianus) [b]
BLUE-BLACK GROSBEAK (Cyanocompsa cyanoides)
Icteridae (Troupials and Allies)
RED-WINGED BLACKBIRD (Agelaius phoeniceus)
RED-BREASTED BLACKBIRD (Sturnella militaris)
EASTERN MEADOWLARK (Sturnella magna)
MELODIOUS BLACKBIRD (Dives dives)
GREAT-TAILED GRACKLE (Quiscalus mexicanus)
NICARAGUAN GRACKLE (Quiscalus nicaraguensis)
BRONZED COWBIRD (Molothrus aeneus)
GIANT COWBIRD (Molothrus oryzivorus)
BLACK-COWLED ORIOLE (Icterus prosthemelas)
ORCHARD ORIOLE (Icterus spurius) [b]
STREAK-BACKED ORIOLE (Icterus pustulatus)
BALTIMORE ORIOLE (Icterus galbula) [b]
YELLOW-BILLED CACIQUE (Amblycercus holosericeus)
SCARLET-RUMPED CACIQUE (SCARLET-RUMPED) (Cacicus uropygialis microrhynchus) [*]
CRESTED OROPENDOLA (Psarocolius decumanus) [N]
MONTEZUMA OROPENDOLA (Psarocolius montezuma)
Fringillidae (Siskins, Crossbills, and Allies)
SCRUB EUPHONIA (Euphonia affinis)
YELLOW-CROWNED EUPHONIA (Euphonia luteicapilla)
THICK-BILLED EUPHONIA (Euphonia laniirostris)
YELLOW-THROATED EUPHONIA (Euphonia hirundinacea)
SPOT-CROWNED EUPHONIA (Euphonia imitans)
OLIVE-BACKED EUPHONIA (Euphonia gouldi)
TAWNY-CAPPED EUPHONIA (Euphonia anneae)
GOLDEN-BROWED CHLOROPHONIA (Chlorophonia callophrys)
LESSER GOLDFINCH (Spinus psaltria)
YELLOW-BELLIED SISKIN (Spinus xanthogastrus)
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus) [I]
CENTRAL AMERICAN WOOLY OPOSSUM (Caluromys derbianus)
LONG-NOSED BAT (Rhynchonycteris naso)
GREATER WHITE-LINED BAT (Saccopteryx bilineata)
GREATER BULLDOG BAT (Noctilio leporinus)
COMMON TENT-MAKING BAT (Uroderma bilobatum)
MANTLED HOWLER MONKEY (Alouatta palliata)
WHITE-THROATED CAPUCHIN (Cebus capucinus)
CENTRAL AMERICAN SPIDER MONKEY (Ateles geoffroyi)
HOFFMANN'S TWO-TOED SLOTH (Choloepus hoffmanni)
VARIEGATED SQUIRREL (Sciurus variegatoides)
RED-TAILED SQUIRREL (Sciurus granatensis)
CENTRAL AMERICAN AGOUTI (Dasyprocta punctata)
PACA (Cuniculus paca)
WHITE-NOSED COATI (Nasua narica)
NEOTROPICAL OTTER (Lontra longicaudis)
Herps
GREEN IGUANA (Iguana iguana)
BLACK SPINY-TAILED IGUANA (Ctenosaura similis)
COMMON BASILISK (Basiliscus basiliscus)
TROPICAL HOUSE GECKO (Hemidactylus mabouia)
CENTRAL AMERICAN WHIPTAIL (Ameiva festiva)
GREEN SPINY LIZARD (Sceloporus malachiticus)
TIGER RAT SNAKE (Spilates pullatus)
FER-DE-LANCE (Bothrops asper)
AMERICAN CROCODILE (Crocodylus acutus)
SPECTACLED CAIMAN (Caiman crocodilus)
CANE TOAD (Bufo marinus)
SMOOTH-SKINNED TOAD (Bufo haematicus)
SMOKY JUNGLE FROG (Leptodactylus pentadactylus)
MESO-AMERICAN SLIDER (Trachemys venusta)
Totals for the tour: 509 bird taxa and 15 mammal taxa