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See this triplist in printable PDF format with media only on page 1.

Fantastic looks at the rare and endangered Marvelous Spatuletail was clearly a tour highlight. (Photo by participant Kevin Heffernan)
Great Trip! But let us not avoid the truth, which is that it could have been greater. The landslide that caused a two-day drive around the landslide and removed three days from the itinerary was a major disappointment. In terms of endemic birds, the landslide could have been in a worse place, but we certainly missed some birds and some scenery while experiencing a bad case of "butt numbification." Still, we managed to capture the last four days of the itinerary in fine fashion and, as this triplist shows, saw a terrific variety of birds including a very high percentage of the specialties, and had many aspects of the tour go very, very well.
Our rapid visit to the Lomas de Lachay before our flight to Chiclayo was aided by some cooperative birds, especially the Thick-billed, Grayish, and Coastal miners and the Cactus Canastero, with the bonus of enough flowers to attract Oasis and other hummingbirds.
After reaching Chiclayo our first destination was Bosque de Pomac (= Batan Grande), where we had to struggle a little to find our first Peruvian Plantcutter and Rufous Flycatcher, but we did, and we added a decent look at the rare Tumbes Swallow, along with many Tumbesian species that were new to us. On the coast at Puerto Eten we had a nice walk on the beach, although the river mouth not only did not have Peruvian Tern -- it had almost no terns or gulls. Before reaching Chaparri, a stop at Tinajones Reservoir was productive, including two Comb Ducks.
Our morning at Chaparri was fun, with tame Sechuran Foxes around the buildings and White-winged Guans throughout the canyon. White-tailed Jays and many other birds sought scraps, and walking up and down the canyon was "Tumbes" territory -- Hummingbird, Tyrant, and Sparrow, along with White-headed Brush-Finches, flocks of Sulphur-throated Finches and Parrot-billed Seedeaters, and Andean Condors and King Vultures overhead.
Our first night of camping preceded another White-winged Guan experience in the dramatic setting of Quebrada Frejolillo (=Limon), where we saw a West Peruvian Screech-Owl by night and a host of Tumbesian species by day, including Ecuadorian Piculet, Black-capped Sparrow, and Gray-and-gold Warbler. After a night in Olmos, we pursued higher-elevation Tumbesian species below Abra Porcuya (= Porculla), finding several difficult species -- Piura Chat-Tyrant, Rufous-necked and Henna-hooded foliage-gleaners -- and some easier ones -- Plumbeous-backed Thrush, Line-cheeked Spinetail, and Black-cowled Saltator.
Based out of Jaen, we birded in arid to moist habitats in the valley of the Rio Chinchipe, finding both more Tumbesian species and some isolates of more widespread South American birds. Highlights included Scrub Nightjar, Maranon Crescentchest, Maranon Spinetail, "Chinchipe" Necklaced Spinetail, and then Little Inca-Finch as we headed toward the Eastern Andes.
Our luck was good at the Huembo Spatuletail Center, where the feeders were hosting multiple Marvelous Spatuletails, which we spent several hours enjoying. After a night at the mystery hotel (now you know what we are talking about, but explaining it is another matter), we were guided up the steep trail above San Lorenzo, both groups managing to see Pale-billed Antpitta along with some other interesting birds, such as Russet-mantled Softtail, Unstreaked Tit-Tyrant, and Plain-tailed Wren.
Based for four nights at Owlet Lodge at Abra Patricia, we moved up and down the east slope from 1000-2300m. The Long-whiskered Owlet was heard at length and glimpsed. We did better in the stunted forest, finding Royal Sunangel and Bar-winged Wood-Wren, while work inside the forest produced views for many of Rusty-tinged and Ochre-fronted antpittas. The hummingbird feeders at the lodge were fun, with a bonus Rufous-capped Thornbill on top of the expected Emerald-bellied Pufflegs and friends (and foes!). Highlights were many, including Lanceolated Monklet, Black-mandibled Toucan, Speckle-chested Piculet, Chestnut-crested Cotinga, Johnson's Tody-Tyrant, and White-capped, Yellow-scarfed, Vermilion, and Blue-browed tanagers. However, flock activity was less than normal, and it was a quieter time than is often the case, the one area on the trip where the birding faltered.
Our long drive was enlivened by only a little birding, with a Green-backed Becard and repeat Peruvian Sheartail, Peruvian Plantcutter, Rufous Flycatcher, and Cinereous Finch standing out. But we were then ready for action from Celendin to Balsas and back, seeing almost everything we sought, particularly Yellow-faced Parrotlet, Peruvian Pigeon, Chestnut-backed Thornbird, Jelski's Chat-Tyrant, Black-crested Tit-tyrant, Maranon Thrush, Buff-bridled and Gray-winged inca-finches, and Rufous-eared Brush-Finch. The high country from Celendin to Cajamarca featured three canasteros, White-tailed Shrike-Tyrant, and Plain-tailed Warbling-Finch. A final day and a few hours around Cajamarca produced Gray-bellied Comet, Black Metaltail, Striated Earthcreeper, Great Spinetail, and Unicolored Tapaculo, along with some eye candy like Purple-throated Sunangel and Golden-rumped Euphonia.
In addition to being a spectacular avifauna, it is a threatened one. As a rough count, we encountered two Critically Endangered, 10 Endangered, 18 Vulnerable, and 20 Near Threatened species, as evaluated by BirdLife International. We know you have your favorite charities, but if this trip developed a special interest in this region, you can channel money to habitat for species you saw to ECOAN (through the American Bird Conservancy if U.S. tax deductions are relevant) for its Huembo and Abra Patricia reserves, and BirdLife International is working on another Spatuletail project right now.
Our journey would not have been possible without the superb crew from Manu, special thanks to their chief and our chef, Aurelio, along with Rene, Alejandro, and Luis.
Taxonomy follows Clements in large part; we are in the process of a transition to Clements 6.8, which may produce a quirk or two. Additional comments rely on various published papers as well as material in the Handbook of the Birds of the World (HBW) and from the South American Classification Committee (S.A.C.C.) website. Apologies are due the Spanish language because we avoid the orthographic marks that do not translate well across the various computer platforms and programs.
--Richard
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Tinamidae (Tinamous)

We all enjoyed up close and personal looks at the very rare White-winged Guan. (Photo by participant Kevin Heffernan)
HOODED TINAMOU (Nothocercus nigrocapillus) [*] TATAUPA TINAMOU (Crypturellus tataupa inops) [*]
ANDEAN TINAMOU (Nothoprocta pentlandii ambigua) [*]
Anatidae (Ducks, Geese, and Waterfowl)
COMB DUCK (Sarkidiornis melanotos)
YELLOW-BILLED PINTAIL (Anas georgica)
YELLOW-BILLED TEAL (Anas flavirostris)
Cracidae (Guans, Chachalacas, and Curassows)
SPECKLED CHACHALACA (Ortalis guttata) [*]
WHITE-WINGED GUAN (Penelope albipennis) [E]
Podicipedidae (Grebes)
PIED-BILLED GREBE (Podilymbus podiceps)
Ciconiidae (Storks)
WOOD STORK (Mycteria americana)
Sulidae (Boobies and Gannets)
PERUVIAN BOOBY (Sula variegata)
Phalacrocoracidae (Cormorants and Shags)
NEOTROPIC CORMORANT (Phalacrocorax brasilianus)
GUANAY CORMORANT (Phalacrocorax bougainvillii)
Pelecanidae (Pelicans)

The Rio Maranon flows through the huge valley it has carved, forming one of the most important biogeographical barriers in the Andes (and South America). The interior valley is a desert in a rainshadow, while the high Eastern Cordillera rises in the background, the more arid Western Cordillera behind the photo. (Photo by guide Richard Webster)
PERUVIAN PELICAN (Pelecanus thagus) Ardeidae (Herons, Egrets, and Bitterns)
FASCIATED TIGER-HERON (Tigrisoma fasciatum)
COCOI HERON (Ardea cocoi)
GREAT EGRET (Ardea alba)
SNOWY EGRET (Egretta thula)
LITTLE BLUE HERON (Egretta caerulea)
TRICOLORED HERON (Egretta tricolor)
CATTLE EGRET (Bubulcus ibis)
STRIATED HERON (Butorides striata)
BLACK-CROWNED NIGHT-HERON (Nycticorax nycticorax)
Threskiornithidae (Ibises and Spoonbills)
PUNA IBIS (Plegadis ridgwayi)
Cathartidae (New World Vultures)
BLACK VULTURE (Coragyps atratus)
TURKEY VULTURE (Cathartes aura)
ANDEAN CONDOR (Vultur gryphus)
KING VULTURE (Sarcoramphus papa)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus) [b]
Accipitridae (Hawks, Eagles, and Kites)
SHARP-SHINNED HAWK (PLAIN-BREASTED) (Accipiter striatus ventralis)

Although this bird is elegant, it is not the Elegant Crescentchest. It is the Maranon Crescentchest with an extremely limited range. (Photo by participant Brian Stech)
SAVANNA HAWK (Buteogallus meridionalis) ROADSIDE HAWK (Rupornis magnirostris)
HARRIS'S HAWK (Parabuteo unicinctus)
WHITE-RUMPED HAWK (Parabuteo leucorrhous)
VARIABLE HAWK (Geranoaetus polyosoma)
BLACK-CHESTED BUZZARD-EAGLE (Geranoaetus melanoleucus)
BROAD-WINGED HAWK (Buteo platypterus) [b]
Rallidae (Rails, Gallinules, and Coots)
SPOTTED RAIL (Pardirallus maculatus)
BLACKISH RAIL (Pardirallus nigricans)
PLUMBEOUS RAIL (Pardirallus sanguinolentus)
COMMON GALLINULE (Gallinula galeata)
SLATE-COLORED COOT (Fulica ardesiaca)
Burhinidae (Thick-knees)
PERUVIAN THICK-KNEE (Burhinus superciliaris)
Charadriidae (Plovers and Lapwings)
ANDEAN LAPWING (Vanellus resplendens)
BLACK-BELLIED PLOVER (Pluvialis squatarola) [b]

Persistence paid off in searching for the Spotted Rail. We caught up to this bird near Olmos. (Photo by participant Kevin Heffernan)
AMERICAN GOLDEN-PLOVER (Pluvialis dominica) [b] SNOWY PLOVER (HUMBOLDT) (Charadrius nivosus occidentalis)
SEMIPALMATED PLOVER (Charadrius semipalmatus) [b]
KILLDEER (Charadrius vociferus)
Haematopodidae (Oystercatchers)
AMERICAN OYSTERCATCHER (Haematopus palliatus)
BLACKISH OYSTERCATCHER (Haematopus ater)
Recurvirostridae (Stilts and Avocets)
BLACK-NECKED STILT (BLACK-NECKED) (Himantopus mexicanus mexicanus)
Scolopacidae (Sandpipers and Allies)
SPOTTED SANDPIPER (Actitis macularius) [b]
SOLITARY SANDPIPER (Tringa solitaria) [b]
GREATER YELLOWLEGS (Tringa melanoleuca) [b]
WHIMBREL (Numenius phaeopus) [b]
RUDDY TURNSTONE (Arenaria interpres) [b]
SANDERLING (Calidris alba) [b]
SEMIPALMATED SANDPIPER (Calidris pusilla) [b]
WESTERN SANDPIPER (Calidris mauri) [b]
LEAST SANDPIPER (Calidris minutilla) [b]
WILSON'S PHALAROPE (Phalaropus tricolor) [b]
Thinocoridae (Seedsnipes)
LEAST SEEDSNIPE (Thinocorus rumicivorus)
Laridae (Gulls, Terns, and Skimmers)
ANDEAN GULL (Chroicocephalus serranus)
GRAY-HOODED GULL (Chroicocephalus cirrocephalus)
GRAY GULL (Leucophaeus modestus)
FRANKLIN'S GULL (Leucophaeus pipixcan) [b]
BELCHER'S GULL (Larus belcheri)

Long-tailed Slyph -- one of the most elegant hummingbirds. (Photo by participant Kevin Heffernan)
KELP GULL (Larus dominicanus) INCA TERN (Larosterna inca)
COMMON TERN (Sterna hirundo) [b]
ROYAL TERN (Thalasseus maximus) [b]
SANDWICH TERN (Thalasseus sandvicensis) [b]
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
BAND-TAILED PIGEON (Patagioenas fasciata)
PERUVIAN PIGEON (Patagioenas oenops)
PLUMBEOUS PIGEON (Patagioenas plumbea)
RUDDY PIGEON (Patagioenas subvinacea) [*]
WEST PERUVIAN DOVE (Zenaida meloda)
EARED DOVE (Zenaida auriculata)
CROAKING GROUND-DOVE (Columbina cruziana)
BLUE GROUND-DOVE (Claravis pretiosa)
BARE-FACED GROUND-DOVE (Metriopelia ceciliae)
WHITE-TIPPED DOVE (Leptotila verreauxi)
WHITE-THROATED QUAIL-DOVE (Geotrygon frenata) [*]
Cuculidae (Cuckoos)
SQUIRREL CUCKOO (Piaya cayana)
STRIPED CUCKOO (Tapera naevia)
SMOOTH-BILLED ANI (Crotophaga ani)
GROOVE-BILLED ANI (Crotophaga sulcirostris)
Strigidae (Owls)
PERUVIAN SCREECH-OWL (Megascops roboratus)
CINNAMON SCREECH-OWL (Megascops petersoni) [*]

Thankfully, Peruvian Pygmy-Owl isn't too hard to find west of the Andes. (Photo by participant Kevin Heffernan)
GREAT HORNED OWL (Bubo virginianus) [*] FERRUGINOUS PYGMY-OWL (Glaucidium brasilianum) [*]
PERUVIAN PYGMY-OWL (Glaucidium peruanum)
LONG-WHISKERED OWLET (Xenoglaux loweryi) [E*]
BURROWING OWL (Athene cunicularia)
RUFOUS-BANDED OWL (Ciccaba albitarsis) [*]
Caprimulgidae (Nightjars and Allies)
LESSER NIGHTHAWK (Chordeiles acutipennis)
BAND-WINGED NIGHTJAR (Caprimulgus longirostris) [*]
SCRUB NIGHTJAR (Caprimulgus anthonyi)
LYRE-TAILED NIGHTJAR (Uropsalis lyra)
Nyctibiidae (Potoos)
COMMON POTOO (Nyctibius griseus) [*]
Apodidae (Swifts)
CHESTNUT-COLLARED SWIFT (Streptoprocne rutila)
WHITE-COLLARED SWIFT (Streptoprocne zonaris)
SHORT-TAILED SWIFT (TUMBES) (Chaetura brachyura ocypetes)

Lyre-tailed Nightjar is one of many species whose range is limited to a narrow elevational band along the Andes. (Photo by participant Kevin Heffernan)
WHITE-TIPPED SWIFT (Aeronautes montivagus) ANDEAN SWIFT (Aeronautes andecolus)
Trochilidae (Hummingbirds)
GRAY-CHINNED HERMIT (Phaethornis griseogularis porcullae)
BROWN VIOLETEAR (Colibri delphinae)
GREEN VIOLETEAR (Colibri thalassinus)
SPARKLING VIOLETEAR (Colibri coruscans)
AMETHYST-THROATED SUNANGEL (Heliangelus amethysticollis)
PURPLE-THROATED SUNANGEL (Heliangelus viola)
ROYAL SUNANGEL (Heliangelus regalis)
SPECKLED HUMMINGBIRD (Adelomyia melanogenys)
LONG-TAILED SYLPH (Aglaiocercus kingi)
GRAY-BELLIED COMET (Taphrolesbia griseiventris) [E]
BLACK-TAILED TRAINBEARER (Lesbia victoriae)

In the right light, the Emerald-bellied Puffleg practically sparkles. (Photo by participant Brian Stech)
GREEN-TAILED TRAINBEARER (Lesbia nuna) RUFOUS-CAPPED THORNBILL (Chalcostigma ruficeps)
TYRIAN METALTAIL (Metallura tyrianthina)
BLACK METALTAIL (Metallura phoebe) [E]
EMERALD-BELLIED PUFFLEG (Eriocnemis aline)
MARVELOUS SPATULETAIL (Loddigesia mirabilis) [E]
SHINING SUNBEAM (Aglaeactis cupripennis) [N]
BRONZY INCA (Coeligena coeligena)
COLLARED INCA (Coeligena torquata)
VIOLET-THROATED STARFRONTLET (Coeligena violifer)
RAINBOW STARFRONTLET (Coeligena iris)
SWORD-BILLED HUMMINGBIRD (Ensifera ensifera)
CHESTNUT-BREASTED CORONET (Boissonneaua matthewsii)
FAWN-BREASTED BRILLIANT (Heliodoxa rubinoides)
VIOLET-FRONTED BRILLIANT (Heliodoxa leadbeateri)
GIANT HUMMINGBIRD (Patagona gigas)
PURPLE-COLLARED WOODSTAR (Myrtis fanny)
OASIS HUMMINGBIRD (Rhodopis vesper)
PERUVIAN SHEARTAIL (Thaumastura cora)
WHITE-BELLIED WOODSTAR (Chaetocercus mulsant)
LITTLE WOODSTAR (Chaetocercus bombus)
SHORT-TAILED WOODSTAR (Myrmia micrura)
FORK-TAILED WOODNYMPH (Thalurania furcata)

Participant Kevin Heffernan shared this beautiful capture of a Green-tailed Trainbearer.
MANY-SPOTTED HUMMINGBIRD (Taphrospilus hypostictus) TUMBES HUMMINGBIRD (Leucippus baeri)
SPOT-THROATED HUMMINGBIRD (Leucippus taczanowskii) [E]
WHITE-BELLIED HUMMINGBIRD (Amazilia chionogaster)
AMAZILIA HUMMINGBIRD (Amazilia amazilia)
ANDEAN EMERALD (Amazilia franciae)
Trogonidae (Trogons)
GOLDEN-HEADED QUETZAL (Pharomachrus auriceps)
CRESTED QUETZAL (Pharomachrus antisianus)
MASKED TROGON (Trogon personatus)
Momotidae (Motmots)
WHOOPING MOTMOT (ARGENTICINCTUS) (Momotus subrufescens argenticinctus)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)
Bucconidae (Puffbirds)
LANCEOLATED MONKLET (Micromonacha lanceolata)
WHITE-FRONTED NUNBIRD (Monasa morphoeus)
Capitonidae (New World Barbets)
VERSICOLORED BARBET (Eubucco versicolor) [*]
Ramphastidae (Toucans)
EMERALD TOUCANET (BLACK-THROATED) (Aulacorhynchus prasinus cyanolaemus)
CHESTNUT-EARED ARACARI (Pteroglossus castanotis)
BLACK-MANDIBLED TOUCAN (Ramphastos ambiguus)
Picidae (Woodpeckers)
ECUADORIAN PICULET (Picumnus sclateri)
SPECKLE-CHESTED PICULET (Picumnus steindachneri) [E]

All of the world's Marvelous Spatuletails live in the valley of the Rio Utcubamba, the main valley of which runs in front of the back ridge. This photograph was taken from the Huembo Spatuletail Center, where we enjoyed great looks at it. (Photo by guide Richard Webster)
YELLOW-TUFTED WOODPECKER (Melanerpes cruentatus) [*] SMOKY-BROWN WOODPECKER (Picoides fumigatus)
LITTLE WOODPECKER (Veniliornis passerinus)
SCARLET-BACKED WOODPECKER (Veniliornis callonotus)
GOLDEN-OLIVE WOODPECKER (Colaptes rubiginosus)
CRIMSON-MANTLED WOODPECKER (Colaptes rivolii)
BLACK-NECKED WOODPECKER (Colaptes atricollis) [E]
SPOT-BREASTED WOODPECKER (Colaptes punctigula)
ANDEAN FLICKER (Colaptes rupicola)
LINEATED WOODPECKER (Dryocopus lineatus)
Falconidae (Falcons and Caracaras)
MOUNTAIN CARACARA (Phalcoboenus megalopterus)
CRESTED CARACARA (Caracara cheriway)
LAUGHING FALCON (Herpetotheres cachinnans)
AMERICAN KESTREL (Falco sparverius)
MERLIN (Falco columbarius) [b]
PEREGRINE FALCON (Falco peregrinus)
Psittacidae (Parrots)
MOUNTAIN PARAKEET (Psilopsiagon aurifrons) [*]

More than a dozen subspecies are recognized for the wide-ranging Golden-olive Woodpecker. (Photo by participant Kevin Heffernan)
SCARLET-FRONTED PARAKEET (Aratinga wagleri) MITRED PARAKEET (Aratinga mitrata) [*]
RED-MASKED PARAKEET (Aratinga erythrogenys)
WHITE-EYED PARAKEET (Aratinga leucophthalma)
BARRED PARAKEET (Bolborhynchus lineola) [*]
PACIFIC PARROTLET (Forpus coelestis)
YELLOW-FACED PARROTLET (Forpus xanthops) [E]
RED-BILLED PARROT (Pionus sordidus)
SCALY-NAPED PARROT (Amazona mercenarius)
Thamnophilidae (Typical Antbirds)
GREAT ANTSHRIKE (Taraba major)
RUFOUS-CAPPED ANTSHRIKE (Thamnophilus ruficapillus jaczewskii)
CHAPMAN'S ANTSHRIKE (Thamnophilus zarumae)
LINED ANTSHRIKE (Thamnophilus tenuepunctatus)
COLLARED ANTSHRIKE (Thamnophilus bernardi)
COLLARED ANTSHRIKE (SHUMBAE) (Thamnophilus bernardi shumbae)
NORTHERN SLATY-ANTSHRIKE (MARAON) (Thamnophilus punctatus leucogaster)
VARIABLE ANTSHRIKE (Thamnophilus caerulescens subandinus)
RUSSET ANTSHRIKE (Thamnistes anabatinus)
SLATY ANTWREN (Myrmotherula schisticolor)
YELLOW-BREASTED ANTWREN (Herpsilochmus axillaris) [*]
LONG-TAILED ANTBIRD (Drymophila caudata)

The beautiful White-tailed Jay is another Tumbesian specialty. (Photo by participant Brian Stech)
BLACKISH ANTBIRD (Cercomacra nigrescens) WHITE-BACKED FIRE-EYE (Pyriglena leuconota) [*]
WHITE-BROWED ANTBIRD (Myrmoborus leucophrys) [*]
COMMON SCALE-BACKED ANTBIRD (Willisornis poecilinotus) [*]
Melanopareiidae (Crescentchests)
MARAON CRESCENTCHEST (Melanopareia maranonica)
ELEGANT CRESCENTCHEST (Melanopareia elegans)
Grallariidae (Antpittas)
CHESTNUT-CROWNED ANTPITTA (Grallaria ruficapilla)
PALE-BILLED ANTPITTA (Grallaria carrikeri) [E]
RUSTY-TINGED ANTPITTA (Grallaria przewalskii) [E]
RUFOUS ANTPITTA (Grallaria rufula cajamarcae)
CHESTNUT ANTPITTA (Grallaria blakei) [E*]
OCHRE-FRONTED ANTPITTA (Grallaricula ochraceifrons) [E]
RUSTY-BREASTED ANTPITTA (TUMBES) (Grallaricula ferrugineipectus leymebambae) [*]
Rhinocryptidae (Tapaculos)
UNICOLORED TAPACULO (Scytalopus unicolor) [E]
TRILLING TAPACULO (Scytalopus parvirostris)
RUFOUS-VENTED TAPACULO (Scytalopus femoralis) [E]
WHITE-CROWNED TAPACULO (Scytalopus atratus atratus)
Formicariidae (Antthrushes)
BARRED ANTTHRUSH (Chamaeza mollissima) [*]
Furnariidae (Ovenbirds and Woodcreepers)
COASTAL MINER (Geositta peruviana) [E]

Unstreaked Tit-Tyrant was an unexpected bonus from our Pale-billed Antpitta hike. (Photo by participant Kevin Heffernan)
SLENDER-BILLED MINER (Geositta tenuirostris) GRAYISH MINER (Geositta maritima)
THICK-BILLED MINER (Geositta crassirostris) [E]
TYRANNINE WOODCREEPER (Dendrocincla tyrannina)
STRONG-BILLED WOODCREEPER (Xiphocolaptes promeropirhynchus)
OLIVE-BACKED WOODCREEPER (Xiphorhynchus triangularis)
STREAK-HEADED WOODCREEPER (Lepidocolaptes souleyetii)
MONTANE WOODCREEPER (Lepidocolaptes lacrymiger)
STREAKED XENOPS (Xenops rutilans)
STREAKED TUFTEDCHEEK (Pseudocolaptes boissonneautii)
PALE-LEGGED HORNERO (PACIFIC) (Furnarius leucopus cinnamomeus)
WREN-LIKE RUSHBIRD (Phleocryptes melanops)
STRIATED EARTHCREEPER (Geocerthia serrana) [E]
CREAM-WINGED CINCLODES (Cinclodes albiventris)
WHITE-WINGED CINCLODES (Cinclodes atacamensis)
SURF CINCLODES (Cinclodes taczanowskii) [E]
BUFF-FRONTED FOLIAGE-GLEANER (Philydor rufum)
MONTANE FOLIAGE-GLEANER (Anabacerthia striaticollis) [N]
BUFF-BROWED FOLIAGE-GLEANER (Syndactyla rufosuperciliata)

Our lunch table at Quebrada Frejolillo, in the shade of a sprawling pink-flowering bougainvilla, a native here. It was attracting a Short-tailed Woodstar, a surprise to us that these flowers attracted any hummingbird. It was a relaxing lunch after a long morning walk, a successful one. (Photo by guide Richard Webster)
RUFOUS-NECKED FOLIAGE-GLEANER (Syndactyla ruficollis) HENNA-HOODED FOLIAGE-GLEANER (Hylocryptus erythrocephalus)
SPOTTED BARBTAIL (Premnoplex brunnescens) [*]
PEARLED TREERUNNER (Margarornis squamiger)
RUSTY-CROWNED TIT-SPINETAIL (Leptasthenura pileata cajabambae) [E]
RUFOUS-FRONTED THORNBIRD (RUFOUS-FRONTED) (Phacellodomus rufifrons peruvianus)
CHESTNUT-BACKED THORNBIRD (Phacellodomus dorsalis) [E]
MANY-STRIPED CANASTERO (Asthenes flammulata)
STREAK-BACKED CANASTERO (Asthenes wyatti)
STREAK-THROATED CANASTERO (Asthenes humilis)
RUSSET-MANTLED SOFTTAIL (Thripophaga berlepschi) [E]
LINE-CHEEKED SPINETAIL (Cranioleuca antisiensis)
BARON'S SPINETAIL (Cranioleuca baroni) [E]
CACTUS CANASTERO (Pseudasthenes cactorum) [E]
AZARA'S SPINETAIL (Synallaxis azarae)
DARK-BREASTED SPINETAIL (Synallaxis albigularis)

Yellow-faced Parrotlet is not the favorite bird of the guides, who know that the search can be short or long, and not always successful--one is looking for 1/1000th of the population. After an unsuccessful afternoon search, our morning started off just fine with several of this attractive parrotlet above Balsas. (Photo by guide Richard Webster)
RUFOUS SPINETAIL (Synallaxis unirufa) MARAON SPINETAIL (Synallaxis maranonica)
NECKLACED SPINETAIL (NECKLACED) (Synallaxis stictothorax maculata)
NECKLACED SPINETAIL (CHINCHIPE) (Synallaxis stictothorax chinchipensis)
GREAT SPINETAIL (Siptornopsis hypochondriaca) [E]
Tyrannidae (Tyrant Flycatchers)
SOUTHERN BEARDLESS-TYRANNULET (Camptostoma obsoletum)
WHITE-BANDED TYRANNULET (Mecocerculus stictopterus)
SULPHUR-BELLIED TYRANNULET (Mecocerculus minor)
BLACK-CRESTED TIT-TYRANT (Anairetes nigrocristatus)
TUFTED TIT-TYRANT (Anairetes parulus)
UNSTREAKED TIT-TYRANT (Anairetes agraphia) [E]
MOUSE-COLORED TYRANNULET (TUMBES) (Phaeomyias murina tumbezana)

Participant Brian Stech caught the light perfectly to showcase the flashy Purple-collared Woodstar.
PACIFIC ELAENIA (Myiopagis subplacens) YELLOW-BELLIED ELAENIA (Elaenia flavogaster)
WHITE-CRESTED ELAENIA (Elaenia albiceps)
MOTTLE-BACKED ELAENIA (Elaenia gigas)
SIERRAN ELAENIA (Elaenia pallatangae)
TORRENT TYRANNULET (Serpophaga cinerea)
STREAK-NECKED FLYCATCHER (Mionectes striaticollis)
OLIVE-STRIPED FLYCATCHER (Mionectes olivaceus)
SLATY-CAPPED FLYCATCHER (Leptopogon superciliaris)
ECUADORIAN TYRANNULET (Phylloscartes gualaquizae)
PERUVIAN TYRANNULET (Zimmerius viridiflavus) [E]
MANY-COLORED RUSH TYRANT (Tachuris rubrigastra)
RUFOUS-HEADED PYGMY-TYRANT (Pseudotriccus ruficeps)
TAWNY-CROWNED PYGMY-TYRANT (Euscarthmus meloryphus)
GRAY-AND-WHITE TYRANNULET (Pseudelaenia leucospodia)
SCALE-CRESTED PYGMY-TYRANT (Lophotriccus pileatus)
BLACK-THROATED TODY-TYRANT (Hemitriccus granadensis)
JOHNSON'S TODY-FLYCATCHER (Poecilotriccus luluae) [E]
COMMON TODY-FLYCATCHER (Todirostrum cinereum)

Rufous Flycatcher is considered "Endangered," with a population under 7,000 birds. Genetically and in plumage it is the outlier in the genus Myiarchus, and may well be split as its own genus, Muscifur. (Photo by guide Richard Webster)
YELLOW-OLIVE FLYCATCHER (Tolmomyias sulphurescens) CINNAMON FLYCATCHER (Pyrrhomyias cinnamomeus)
CLIFF FLYCATCHER (Hirundinea ferruginea)
OLIVE-CHESTED FLYCATCHER (Myiophobus cryptoxanthus)
BRAN-COLORED FLYCATCHER (Myiophobus fasciatus)
WESTERN WOOD-PEWEE (Contopus sordidulus) [b]
TROPICAL PEWEE (TUMBES) (Contopus cinereus punensis)
BLACK PHOEBE (Sayornis nigricans)
VERMILION FLYCATCHER (Pyrocephalus rubinus)
RUFOUS-TAILED TYRANT (Knipolegus poecilurus)
WHITE-WINGED BLACK-TYRANT (WHITE-WINGED) (Knipolegus aterrimus heterogyna)
RUFOUS-NAPED GROUND-TYRANT (Muscisaxicola rufivertex)
BLACK-BILLED SHRIKE-TYRANT (Agriornis montanus)
WHITE-TAILED SHRIKE-TYRANT (Agriornis albicauda)
STREAK-THROATED BUSH-TYRANT (Myiotheretes striaticollis)
RUFOUS-WEBBED BUSH-TYRANT (Polioxolmis rufipennis)
TUMBES TYRANT (Tumbezia salvini)

Celendin, at an elevation of 8,600' (2600m) is a convenient base between the Maranon at Balsas and Cajamarca. Our breakfast birding stop had included Rufous-eared Brush-Finch and White-tailed Shrike-Tyrant, and we were headed for Gray-winged Inca-Finches. (Photo by guide Richard Webster)
JELSKI'S CHAT-TYRANT (Ochthoeca jelskii) RUFOUS-BREASTED CHAT-TYRANT (Ochthoeca rufipectoralis)
PIURA CHAT-TYRANT (Ochthoeca piurae) [E]
WHITE-BROWED CHAT-TYRANT (Ochthoeca leucophrys)
LONG-TAILED TYRANT (Colonia colonus)
SHORT-TAILED FIELD TYRANT (Muscigralla brevicauda)
RUFOUS FLYCATCHER (Myiarchus semirufus) [E]
DUSKY-CAPPED FLYCATCHER (Myiarchus tuberculifer atriceps)
SOOTY-CROWNED FLYCATCHER (Myiarchus phaeocephalus phaeocephalus)
PALE-EDGED FLYCATCHER (Myiarchus cephalotes)
BROWN-CRESTED FLYCATCHER (Myiarchus tyrannulus)
GREAT KISKADEE (Pitangus sulphuratus)
SOCIAL FLYCATCHER (Myiozetetes similis)
GOLDEN-CROWNED FLYCATCHER (Myiodynastes chrysocephalus)
BAIRD'S FLYCATCHER (Myiodynastes bairdii)
PIRATIC FLYCATCHER (Legatus leucophaius)
TROPICAL KINGBIRD (Tyrannus melancholicus)
EASTERN KINGBIRD (Tyrannus tyrannus) [b]
Cotingidae (Cotingas)
GREEN-AND-BLACK FRUITEATER (Pipreola riefferii)
PERUVIAN PLANTCUTTER (Phytotoma raimondii) [E]
RED-CRESTED COTINGA (Ampelion rubrocristatus)
CHESTNUT-CRESTED COTINGA (Ampelion rufaxilla)
ANDEAN COCK-OF-THE-ROCK (Rupicola peruvianus)
Tityridae (Tityras and Allies)

Vermilion Flycatcher -- a common species but one you never tire of seeing. (Photo by participant Kevin Heffernan)
GREEN-BACKED BECARD (YELLOW-CHEEKED) (Pachyramphus viridis xanthogenys) BARRED BECARD (Pachyramphus versicolor)
BLACK-AND-WHITE BECARD (Pachyramphus albogriseus)
Vireonidae (Vireos)
RED-EYED VIREO (Vireo olivaceus)
OLIVACEOUS GREENLET (Hylophilus olivaceus)
RUFOUS-BROWED PEPPERSHRIKE (Cyclarhis gujanensis)
Corvidae (Crows, Jays, and Magpies)
GREEN JAY (Cyanocorax yncas)
WHITE-TAILED JAY (Cyanocorax mystacalis)
Hirundinidae (Swallows)
BLUE-AND-WHITE SWALLOW (Pygochelidon cyanoleuca)
BROWN-BELLIED SWALLOW (Orochelidon murina)
WHITE-BANDED SWALLOW (Atticora fasciata)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)
GRAY-BREASTED MARTIN (Progne chalybea)
TUMBES SWALLOW (Tachycineta stolzmanni)
WHITE-WINGED SWALLOW (Tachycineta albiventer)
BARN SWALLOW (Hirundo rustica) [b]
CHESTNUT-COLLARED SWALLOW (Petrochelidon rufocollaris)
Troglodytidae (Wrens)
SCALY-BREASTED WREN (Microcerculus marginatus) [*]
GRAY-MANTLED WREN (Odontorchilus branickii)
HOUSE WREN (Troglodytes aedon)
MOUNTAIN WREN (Troglodytes solstitialis)
SEDGE WREN (Cistothorus platensis)
FASCIATED WREN (Campylorhynchus fasciatus)
THRUSH-LIKE WREN (Campylorhynchus turdinus)
PLAIN-TAILED WREN (Pheugopedius euophrys schulenbergi)
SPECKLE-BREASTED WREN (SPECKLE-BREASTED) (Pheugopedius sclateri paucimaculatus)
SPECKLE-BREASTED WREN (MARAON) (Pheugopedius sclateri sclateri)

The presence of Golden-billed Saltators is often first detected by their loud song. (Photo by participant Kevin Heffernan)
SUPERCILIATED WREN (Cantorchilus superciliaris) SHARPE'S WREN (Cinnycerthia olivascens)
PERUVIAN WREN (Cinnycerthia peruana) [*]
BAR-WINGED WOOD-WREN (Henicorhina leucoptera)
GRAY-BREASTED WOOD-WREN (Henicorhina leucophrys)
CHESTNUT-BREASTED WREN (Cyphorhinus thoracicus) [*]
Polioptilidae (Gnatcatchers)
TROPICAL GNATCATCHER (WHITE-BROWED) (Polioptila plumbea bilineata)
TROPICAL GNATCATCHER (MARAON) (Polioptila plumbea maior)
Donacobiidae (Donacobius)
BLACK-CAPPED DONACOBIUS (Donacobius atricapilla)
Turdidae (Thrushes and Allies)
ANDEAN SOLITAIRE (Myadestes ralloides)
SLATY-BACKED NIGHTINGALE-THRUSH (Catharus fuscater) [*]
SWAINSON'S THRUSH (Catharus ustulatus) [b]
PLUMBEOUS-BACKED THRUSH (Turdus reevei)
BLACK-BILLED THRUSH (Turdus ignobilis)
MARAON THRUSH (Turdus maranonicus)
GREAT THRUSH (Turdus fuscater)
CHIGUANCO THRUSH (Turdus chiguanco)
GLOSSY-BLACK THRUSH (Turdus serranus)
Mimidae (Mockingbirds and Thrashers)
LONG-TAILED MOCKINGBIRD (Mimus longicaudatus)
Motacillidae (Wagtails and Pipits)

The dramatic landscape of Quebrada Frejolillo, where we sought White-winged Guans in one of their most important centers of breeding, and otherwise enjoyed a rich selection of Tumbesian birds, including Black-capped Sparrow and Gray-and-gold Warbler. (Photo by guide Richard Webster)
PARAMO PIPIT (Anthus bogotensis) Parulidae (New World Warblers)
MASKED YELLOWTHROAT (MASKED) (Geothlypis aequinoctialis peruviana)
MASKED YELLOWTHROAT (BLACK-LORED) (Geothlypis aequinoctialis auricularis)
TROPICAL PARULA (Setophaga pitiayumi)
BLACKBURNIAN WARBLER (Setophaga fusca) [b]
THREE-STRIPED WARBLER (Basileuterus tristriatus)
THREE-BANDED WARBLER (Basileuterus trifasciatus)
CITRINE WARBLER (Myiothlypis luteoviridis)
BLACK-CRESTED WARBLER (Myiothlypis nigrocristata)
GRAY-AND-GOLD WARBLER (Myiothlypis fraseri)
RUSSET-CROWNED WARBLER (Myiothlypis coronata)
CANADA WARBLER (Cardellina canadensis) [b]
SLATE-THROATED REDSTART (Myioborus miniatus)
SPECTACLED REDSTART (Myioborus melanocephalus)
Thraupidae (Tanagers and Allies)
MAGPIE TANAGER (Cissopis leverianus)
WHITE-CAPPED TANAGER (Sericossypha albocristata)
RUFOUS-CRESTED TANAGER (Creurgops verticalis)
BLACK-CAPPED HEMISPINGUS (WHITE-BROWED) (Hemispingus atropileus auricularis)
DRAB HEMISPINGUS (Hemispingus xanthophthalmus)
GRAY-HOODED BUSH TANAGER (Cnemoscopus rubrirostris)
RUFOUS-CHESTED TANAGER (Thlypopsis ornata)

Three-banded Warbler -- a Tumbesian specialty. (Photo by participant Kevin Heffernan)
BUFF-BELLIED TANAGER (Thlypopsis inornata) WHITE-LINED TANAGER (Tachyphonus rufus)
BLACK-BELLIED TANAGER (Ramphocelus melanogaster) [E]
BLUE-GRAY TANAGER (Thraupis episcopus)
PALM TANAGER (Thraupis palmarum)
BLUE-CAPPED TANAGER (Thraupis cyanocephala)
BLUE-AND-YELLOW TANAGER (Thraupis bonariensis darwinii)
VERMILION TANAGER (Calochaetes coccineus)
HOODED MOUNTAIN-TANAGER (Buthraupis montana)
LACRIMOSE MOUNTAIN-TANAGER (Anisognathus lacrymosus)
SCARLET-BELLIED MOUNTAIN-TANAGER (Anisognathus igniventris)
BLUE-WINGED MOUNTAIN-TANAGER (Anisognathus somptuosus)
GRASS-GREEN TANAGER (Chlorornis riefferii)
YELLOW-SCARFED TANAGER (Iridosornis reinhardti) [E]
FAWN-BREASTED TANAGER (Pipraeidea melanonota)
SILVERY TANAGER (Tangara viridicollis)
BLUE-NECKED TANAGER (Tangara cyanicollis)
SPOTTED TANAGER (Tangara punctata)
BLUE-AND-BLACK TANAGER (Tangara vassorii)
BERYL-SPANGLED TANAGER (Tangara nigroviridis)
BLUE-BROWED TANAGER (Tangara cyanotis)
PARADISE TANAGER (Tangara chilensis)
BAY-HEADED TANAGER (Tangara gyrola)
GOLDEN-EARED TANAGER (Tangara chrysotis)
SAFFRON-CROWNED TANAGER (Tangara xanthocephala)
FLAME-FACED TANAGER (Tangara parzudakii)
GREEN-AND-GOLD TANAGER (Tangara schrankii)
GOLDEN TANAGER (Tangara arthus)

We studied a pair of Lineated Woodpeckers at Chaparri. (Photo by participant Kevin Heffernan)
SWALLOW TANAGER (Tersina viridis) BLACK-FACED DACNIS (Dacnis lineata)
BLUE DACNIS (Dacnis cayana)
PURPLE HONEYCREEPER (Cyanerpes caeruleus)
GREEN HONEYCREEPER (Chlorophanes spiza)
CINEREOUS CONEBILL (Conirostrum cinereum)
CAPPED CONEBILL (Conirostrum albifrons)
TIT-LIKE DACNIS (Xenodacnis parina)
BLACK-THROATED FLOWERPIERCER (Diglossa brunneiventris)
WHITE-SIDED FLOWERPIERCER (Diglossa albilatera)
RUSTY FLOWERPIERCER (Diglossa sittoides)
BLUISH FLOWERPIERCER (Diglossa caerulescens)
MASKED FLOWERPIERCER (Diglossa cyanea)
PERUVIAN SIERRA-FINCH (Phrygilus punensis)
MOURNING SIERRA-FINCH (Phrygilus fruticeti)
PLUMBEOUS SIERRA-FINCH (Phrygilus unicolor)
ASH-BREASTED SIERRA-FINCH (Phrygilus plebejus)
CINEREOUS FINCH (Piezorhina cinerea) [E]
GRAY-WINGED INCA-FINCH (Incaspiza ortizi) [E]
BUFF-BRIDLED INCA-FINCH (Incaspiza laeta) [EN]
LITTLE INCA-FINCH (Incaspiza watkinsi) [E]
PLAIN-TAILED WARBLING-FINCH (Poospiza alticola) [E]
COLLARED WARBLING-FINCH (Poospiza hispaniolensis)
SAFFRON FINCH (Sicalis flaveola)
SULPHUR-THROATED FINCH (Sicalis taczanowskii)
BLUE-BLACK GRASSQUIT (Volatinia jacarina)
BLACK-AND-WHITE SEEDEATER (Sporophila luctuosa)
YELLOW-BELLIED SEEDEATER (Sporophila nigricollis)
PARROT-BILLED SEEDEATER (Sporophila peruviana)
DRAB SEEDEATER (Sporophila simplex)
CHESTNUT-THROATED SEEDEATER (Sporophila telasco)

Buff-bridled Inca-Finch are one of the species restricted to the Maranon Valley. (Photo by participant Kevin Heffernan)
BAND-TAILED SEEDEATER (Catamenia analis) PLAIN-COLORED SEEDEATER (Catamenia inornata)
RED-CRESTED FINCH (Coryphospingus cucullatus)
BANANAQUIT (Coereba flaveola)
DULL-COLORED GRASSQUIT (Tiaris obscurus)
STREAKED SALTATOR (Saltator striatipectus immaculatus)
STREAKED SALTATOR (Saltator striatipectus peruvianus)
GRAYISH SALTATOR (Saltator coerulescens)
BLACK-COWLED SALTATOR (Saltator nigriceps)
GOLDEN-BILLED SALTATOR (Saltator aurantiirostris)
BUFF-THROATED SALTATOR (Saltator maximus)
SLATE-COLORED GROSBEAK (Saltator grossus) [*]
Emberizidae (Buntings and New World Sparrows)
BLACK-CAPPED SPARROW (Arremon abeillei abeillei)
BLACK-CAPPED SPARROW (Arremon abeillei nigriceps)
GRAY-BROWED BRUSH-FINCH (Arremon assimilis)
YELLOW-BREASTED BRUSH-FINCH (Atlapetes latinuchus latinuchus)
YELLOW-BREASTED BRUSH-FINCH (Atlapetes latinuchus baroni)
RUFOUS-EARED BRUSH-FINCH (Atlapetes rufigenis) [E]
WHITE-WINGED BRUSH-FINCH (Atlapetes leucopterus)
WHITE-HEADED BRUSH-FINCH (Atlapetes albiceps)
BAY-CROWNED BRUSH-FINCH (Atlapetes seebohmi)
TUMBES SPARROW (Rhynchospiza stolzmanni)
YELLOW-BROWED SPARROW (Ammodramus aurifrons)
RUFOUS-COLLARED SPARROW (Zonotrichia capensis)
COMMON BUSH-TANAGER (NORTHERN ANDES) (Chlorospingus ophthalmicus hiaticolus)
YELLOW-THROATED BUSH-TANAGER (Chlorospingus flavigularis)
ASHY-THROATED BUSH-TANAGER (Chlorospingus canigularis)
Cardinalidae (Cardinals and Allies)

Participant Kevin Heffernan scored this excellent photo of a Collared Inca at the feeding station at the Owlet Lodge.
HEPATIC TANAGER (HIGHLAND) (Piranga flava lutea) SUMMER TANAGER (Piranga rubra) [b]
WHITE-WINGED TANAGER (Piranga leucoptera)
GOLDEN-BELLIED GROSBEAK (Pheucticus chrysogaster)
Icteridae (Troupials and Allies)
PERUVIAN MEADOWLARK (Sturnella bellicosa)
SCRUB BLACKBIRD (Dives warszewiczi)
ORIOLE BLACKBIRD (Gymnomystax mexicanus)
SHINY COWBIRD (Molothrus bonariensis)
GIANT COWBIRD (Molothrus oryzivorus)
WHITE-EDGED ORIOLE (Icterus graceannae)
YELLOW-TAILED ORIOLE (Icterus mesomelas)
MOUNTAIN CACIQUE (GOLDEN-SHOULDERED) (Cacicus chrysonotus peruvianus)
SCARLET-RUMPED CACIQUE (Cacicus uropygialis)
YELLOW-RUMPED CACIQUE (Cacicus cela)
RUSSET-BACKED OROPENDOLA (Psarocolius angustifrons)
CRESTED OROPENDOLA (Psarocolius decumanus)
Fringillidae (Siskins, Crossbills, and Allies)
PURPLE-THROATED EUPHONIA (Euphonia chlorotica)
THICK-BILLED EUPHONIA (Euphonia laniirostris)
GOLDEN-RUMPED EUPHONIA (Euphonia cyanocephala)
BRONZE-GREEN EUPHONIA (Euphonia mesochrysa)
WHITE-VENTED EUPHONIA (Euphonia minuta)
ORANGE-BELLIED EUPHONIA (Euphonia xanthogaster)
LESSER GOLDFINCH (Spinus psaltria)
HOODED SISKIN (Spinus magellanicus)
OLIVACEOUS SISKIN (Spinus olivaceus)
Passeridae (Old World Sparrows)

We arrived at our campsite in Balsas on a lovely evening, the setting sun lighting up the clouds as the Rio Maranon, barely visible in this photograph, flowed quietly (but strongly) through this immense valley. (Photo by guide Richard Webster)
HOUSE SPARROW (Passer domesticus) [I]
GUAYAQUIL SQUIRREL (Sciurus stramineus)
SECHURAN FOX (Pseudalopex sechurae)
TAYRA (Eira barbara)
COLLARED PECCARY (Tayassu tajacu)
WHITE-TAILED DEER (Odocoileus virginianus)
There were, of course, many other critters, including:
Iguana
several smaller lizard species, probably including Dicrodon guttulatum in the coastal deserts (Blue-headed Whiptail) and Microlophus (Tropidurus) koepckeorum above Balsas in the Maranon Valley (Koepcke's Curly-tailed Lizard)
Bat sp. several times
Cicadas and more cicadas
Millipedes and more millipedes
That fascinating Tarantula Hawk/Pepsis Wasp with its spider prey
That less fascinating tarantula that was in someone's shower!
An impressive selection of moths attracted by the lights at Owlet Lodge
Morphos repeatedly, and a moderate selection of other lovely butterflies
white-winged grasshoppers at Aguas Verdes
Totals for the tour: 502 bird taxa and 5 mammal taxa