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The main target of this extension to the Beni -- the rare and beautiful Blue-throated Macaw, here at home in an Atalea palm grove. (Photo by guide Dan Lane)
What is “The Beni” and why is it so durned special? Well, it’s believed that over a million years ago, much of the southwestern Amazon basin was occupied by an inland sea. Over the millennia, as the Andes rose, this sea started to recede, draining out through the proto-Amazon. The last of this sea was left in the present-day Beni, a low-lying area that drained through the bottleneck of the geologic arch on what is the present-day Rio Madeira, just inside Brazil. Today, the Beni still becomes a large lake during the rainy season, but drains for the most part during the dry season, as we saw. While it was still a body of water, it is conjectured that there were nevertheless some higher lands that existed within it as islands. These islands probably were vegetated with a forest reminiscent of ‘varzea’ present today in much of Amazonia, and birds related to species found in Amazonian varzea were isolated on these islands long enough to differentiate. As the Beni “dried out”, an Amazonian influence was able to extend up the gallery forests along the larger rivers that drained it (particularly the Rio Mamore), but others were excluded by the presence of competitors already present in the area. Meanwhile, the floodplain became vegetated by plants that are shared with the Pantanal, an adjacent region that probably shared a similar history (although is not connected directly to the Amazon), resulting in the palm-dominated savanna we see there today.
“Lake Beni” was inhabited by the Amazon River Dolphin, but because of the continued presence of the rocky arch on the Rio Madeira, forming rapids today, the Beni population and that in the Amazon proper have became isolated, and now two taxa exist: species in the eyes of some mammalogists. Similar things happened to several birds such as Plain Softtail, Velvet-fronted Grackle, Varzea and Unicolored thrushes, and a pair of macaws with blue and yellow plumage. The first three are not all that strongly differentiated -- the Amazonian and Beni forms of the first two are, as you can see, still considered conspecifics, but further study is likely to overturn that status. The last pair is more clearly differentiated and, more importantly, are now found together without interbreeding…clear evidence that they cannot be considered conspecifics! Sadly, the present-day co-occurrence of Blue-throated and Blue-and-yellow macaws may be the undoing of the former. The larger and more aggressive Blue-and-yellow Macaw occupies a similar niche to its smaller cousin and, worse still, seems to be capable of muscling it out of breeding sites. The smaller Blue-throated Macaw, to its disadvantage, is also more specialized than its larger relative, requiring large stands of the Atalea palm for foraging and nesting places. It was in such a place that we went to see the pair we enjoyed. Of course, the interest in the rarer macaw by the pet trade hasn’t helped this bird, either. With luck, ecotourism may help bring conservation dollars in to help it withstand these perils and continue to screech amid the Beni's palms.
But the Beni is more than a land with a few specialized birds! As we found, it is in fact a very birdy place, what with all the large waterbirds, the screamers, the limpkins, the storks, the ducks, the skimmers and terns, and then there are the parrots, parakeets, pigeons, ground-doves, finches, horneros, cachalotes, woodpeckers, woodcreepers, spinetails…well, you get the idea! In less than four days, we saw about half the number of species we enjoyed on the following two weeks on the main Bolivia tour! Wow, that’s some bird biomass! And let’s not forget the caimans, capybaras, foxes, and that incredible experience watching the mating tangle of anacondas! Yes sir, the Beni is a pretty special place. I know I enjoyed the visit, and I hope you all did too. Keep those binoculars handy, and maybe see you again soon on another tour!
--Dan
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Rheidae (Rheas)
GREATER RHEA (Rhea americana)
Tinamidae (Tinamous)
UNDULATED TINAMOU (Crypturellus undulatus)
SMALL-BILLED TINAMOU (Crypturellus parvirostris)
RED-WINGED TINAMOU (Rhynchotus rufescens)
Anhimidae (Screamers)
SOUTHERN SCREAMER (Chauna torquata)
Anatidae (Ducks, Geese, and Waterfowl)
WHITE-FACED WHISTLING-DUCK (Dendrocygna viduata)
BLACK-BELLIED WHISTLING-DUCK (Dendrocygna autumnalis)
ORINOCO GOOSE (Neochen jubata)
MUSCOVY DUCK (Cairina moschata)
BRAZILIAN TEAL (Amazonetta brasiliensis)
Cracidae (Guans, Chachalacas, and Curassows)
SPECKLED CHACHALACA (Ortalis guttata)
Ciconiidae (Storks)
MAGUARI STORK (Ciconia maguari)
JABIRU (Jabiru mycteria)
WOOD STORK (Mycteria americana)
Phalacrocoracidae (Cormorants and Shags)
NEOTROPIC CORMORANT (Phalacrocorax brasilianus)
Anhingidae (Anhingas)
ANHINGA (Anhinga anhinga)
Ardeidae (Herons, Egrets, and Bitterns)
RUFESCENT TIGER-HERON (Tigrisoma lineatum)
COCOI HERON (Ardea cocoi)
GREAT EGRET (Ardea alba)
SNOWY EGRET (Egretta thula)
CATTLE EGRET (Bubulcus ibis)
STRIATED HERON (Butorides striata)
WHISTLING HERON (Syrigma sibilatrix)
CAPPED HERON (Pilherodius pileatus)
BLACK-CROWNED NIGHT-HERON (Nycticorax nycticorax)
Threskiornithidae (Ibises and Spoonbills)
GREEN IBIS (Mesembrinibis cayennensis)
BARE-FACED IBIS (Phimosus infuscatus)
PLUMBEOUS IBIS (Theristicus caerulescens)
BUFF-NECKED IBIS (Theristicus caudatus)

As the Beni dries out, birds tend to congregate around the remaining pools of water, as these Wood Storks, Roseate Spoonbills, and Jabirus are doing. (Photo by guide Dan Lane)
ROSEATE SPOONBILL (Platalea ajaja) Cathartidae (New World Vultures)
BLACK VULTURE (Coragyps atratus)
TURKEY VULTURE (Cathartes aura)
LESSER YELLOW-HEADED VULTURE (Cathartes burrovianus)
KING VULTURE (Sarcoramphus papa)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus) [b]
Accipitridae (Hawks, Eagles, and Kites)
SNAIL KITE (Rostrhamus sociabilis)
PLUMBEOUS KITE (Ictinia plumbea)
BLACK-COLLARED HAWK (Busarellus nigricollis)
CRANE HAWK (Geranospiza caerulescens)
GREAT BLACK-HAWK (Buteogallus urubitinga)
SAVANNA HAWK (Buteogallus meridionalis)
ROADSIDE HAWK (Buteo magnirostris)
WHITE-TAILED HAWK (Buteo albicaudatus)
Falconidae (Falcons and Caracaras)
SOUTHERN CARACARA (Caracara plancus)
YELLOW-HEADED CARACARA (Milvago chimachima)
AMERICAN KESTREL (Falco sparverius)
APLOMADO FALCON (Falco femoralis)
Rallidae (Rails, Gallinules, and Coots)
RUFOUS-SIDED CRAKE (Laterallus melanophaius)
GRAY-NECKED WOOD-RAIL (Aramides cajanea)
Aramidae (Limpkin)
LIMPKIN (Aramus guarauna)
Charadriidae (Plovers and Lapwings)
SOUTHERN LAPWING (Vanellus chilensis)
Jacanidae (Jacanas)
WATTLED JACANA (Jacana jacana)
Scolopacidae (Sandpipers and Allies)
SOLITARY SANDPIPER (Tringa solitaria)
LESSER YELLOWLEGS (Tringa flavipes)
UPLAND SANDPIPER (Bartramia longicauda)
WHITE-RUMPED SANDPIPER (Calidris fuscicollis)
PECTORAL SANDPIPER (Calidris melanotos)
STILT SANDPIPER (Calidris himantopus)
Laridae (Gulls, Terns, and Skimmers)
YELLOW-BILLED TERN (Sternula superciliaris)
LARGE-BILLED TERN (Phaetusa simplex)
BLACK SKIMMER (CINERASCENS) (Rynchops niger cinerascens)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia)
PALE-VENTED PIGEON (Patagioenas cayennensis)
PICAZURO PIGEON (Patagioenas picazuro)
EARED DOVE (Zenaida auriculata)
PLAIN-BREASTED GROUND-DOVE (Columbina minuta)
RUDDY GROUND-DOVE (Columbina talpacoti)
PICUI GROUND-DOVE (Columbina picui)
WHITE-TIPPED DOVE (Leptotila verreauxi)
GRAY-FRONTED DOVE (Leptotila rufaxilla)
Psittacidae (Parrots)
MITRED PARAKEET (Aratinga mitrata) [I]
WHITE-EYED PARAKEET (Aratinga leucophthalma)
DUSKY-HEADED PARAKEET (Aratinga weddellii)

An unexpected highlight of the trip was seeing a tangle of Anacondas -- an immense female, seen here, along with three smaller males, all competing to mate with her. (Photo by guide Dan Lane)
PEACH-FRONTED PARAKEET (Aratinga aurea) CHESTNUT-FRONTED MACAW (Ara severus)
RED-AND-GREEN MACAW (Ara chloropterus)
BLUE-AND-YELLOW MACAW (Ara ararauna)
BLUE-THROATED MACAW (Ara glaucogularis) [E]
YELLOW-COLLARED MACAW (Primolius auricollis)
BLUE-WINGED PARROTLET (Forpus xanthopterygius)
YELLOW-CHEVRONED PARAKEET (Brotogeris chiriri)
SCALY-HEADED PARROT (Pionus maximiliani siy)
BLUE-FRONTED PARROT (Amazona aestiva)
Opisthocomidae (Hoatzin)
HOATZIN (Opisthocomus hoazin)
Cuculidae (Cuckoos)
SQUIRREL CUCKOO (Piaya cayana)
GUIRA CUCKOO (Guira guira)
STRIPED CUCKOO (Tapera naevia)
SMOOTH-BILLED ANI (Crotophaga ani)
Strigidae (Owls)
FERRUGINOUS PYGMY-OWL (Glaucidium brasilianum)
BURROWING OWL (Athene cunicularia)
Caprimulgidae (Nightjars and Allies)
NACUNDA NIGHTHAWK (Chordeiles nacunda)
BAND-TAILED NIGHTHAWK (Nyctiprogne leucopyga)
SCISSOR-TAILED NIGHTJAR (Hydropsalis torquata)
Nyctibiidae (Potoos)
GREAT POTOO (Nyctibius grandis)
Apodidae (Swifts)
SHORT-TAILED SWIFT (Chaetura brachyura)
Trochilidae (Hummingbirds)
WHITE-TAILED GOLDENTHROAT (Polytmus guainumbi)
BLACK-THROATED MANGO (Anthracothorax nigricollis)
SWALLOW-TAILED HUMMINGBIRD (Eupetomena macroura)
SAPPHIRE-SPANGLED EMERALD (Amazilia lactea)
GILDED HUMMINGBIRD (Hylocharis chrysura)
Trogonidae (Trogons)
BLACK-TAILED TROGON (Trogon melanurus)
BLUE-CROWNED TROGON (Trogon curucui)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)
AMAZON KINGFISHER (Chloroceryle amazona)
GREEN KINGFISHER (Chloroceryle americana)
AMERICAN PYGMY KINGFISHER (Chloroceryle aenea)
Bucconidae (Puffbirds)
BLACK-FRONTED NUNBIRD (Monasa nigrifrons)
Galbulidae (Jacamars)
RUFOUS-TAILED JACAMAR (Galbula ruficauda)
Ramphastidae (Toucans)
CHESTNUT-EARED ARACARI (Pteroglossus castanotis)
TOCO TOUCAN (Ramphastos toco)
Picidae (Woodpeckers)
WHITE-WEDGED PICULET (Picumnus albosquamatus)
WHITE WOODPECKER (Melanerpes candidus)
YELLOW-TUFTED WOODPECKER (Melanerpes cruentatus)
LITTLE WOODPECKER (Veniliornis passerinus)
GOLDEN-GREEN WOODPECKER (Piculus chrysochloros)
SPOT-BREASTED WOODPECKER (Colaptes punctigula)
GREEN-BARRED WOODPECKER (Colaptes melanochloros)
CAMPO FLICKER (Colaptes campestris)
PALE-CRESTED WOODPECKER (Celeus lugubris)
LINEATED WOODPECKER (Dryocopus lineatus)
CRIMSON-CRESTED WOODPECKER (Campephilus melanoleucos)
Furnariidae (Ovenbirds and Woodcreepers)
RUFOUS HORNERO (Furnarius rufus)
CHOTOY SPINETAIL (Schoeniophylax phryganophilus)
PALE-BREASTED SPINETAIL (Synallaxis albescens australis)
PLAIN-CROWNED SPINETAIL (Synallaxis gujanensis inornata)
RUSTY-BACKED SPINETAIL (Cranioleuca vulpina)
YELLOW-CHINNED SPINETAIL (Certhiaxis cinnamomeus)
PLAIN SOFTTAIL (Thripophaga fusciceps fusciceps)
GREATER THORNBIRD (Phacellodomus ruber)
RUFOUS CACHOLOTE (Pseudoseisura unirufa)
OLIVACEOUS WOODCREEPER (AMAZONIAN) (Sittasomus griseicapillus viridis)
GREAT RUFOUS WOODCREEPER (Xiphocolaptes major)
STRAIGHT-BILLED WOODCREEPER (Dendroplex picus)
BUFF-THROATED WOODCREEPER (Xiphorhynchus guttatus)
NARROW-BILLED WOODCREEPER (Lepidocolaptes angustirostris)
RED-BILLED SCYTHEBILL (Campylorhamphus trochilirostris)
Thamnophilidae (Typical Antbirds)
GREAT ANTSHRIKE (Taraba major)
BARRED ANTSHRIKE (Thamnophilus doliatus)
BOLIVIAN SLATY-ANTSHRIKE (Thamnophilus sticturus)
RUSTY-BACKED ANTWREN (Formicivora rufa)
MATO GROSSO ANTBIRD (Cercomacra melanaria)
BAND-TAILED ANTBIRD (Hypocnemoides maculicauda)
Tyrannidae (Tyrant Flycatchers)
SOUTHERN BEARDLESS-TYRANNULET (Camptostoma obsoletum)

Among the many unexplained mysteries in the bird world is why the name Dull-capped Attila was chosen for this species over its alternate name, White-eyed Attila. (Photo by Guide Dan Lane)
YELLOW TYRANNULET (Capsiempis flaveola) CRESTED DORADITO (Pseudocolopteryx sclateri)
FOREST ELAENIA (Myiopagis gaimardii)
YELLOW-BELLIED ELAENIA (Elaenia flavogaster)
LARGE ELAENIA (Elaenia spectabilis)
WHITE-BELLIED TYRANNULET (Serpophaga munda)
SOUTHERN SCRUB-FLYCATCHER (Sublegatus modestus)
PLAIN TYRANNULET (Inezia inornata)
PEARLY-VENTED TODY-TYRANT (Hemitriccus margaritaceiventer)
RUSTY-FRONTED TODY-FLYCATCHER (Poecilotriccus latirostris)
COMMON TODY-FLYCATCHER (Todirostrum cinereum)
YELLOW-OLIVE FLYCATCHER (MATO GROSSO) (Tolmomyias sulphurescens pallescens)
BRAN-COLORED FLYCATCHER (Myiophobus fasciatus)
VERMILION FLYCATCHER (Pyrocephalus rubinus)
HUDSON'S BLACK-TYRANT (Knipolegus hudsoni)
SPECTACLED TYRANT (Hymenops perspicillatus)
YELLOW-BROWED TYRANT (Satrapa icterophrys)
GRAY MONJITA (Xolmis cinereus)
WHITE-RUMPED MONJITA (Xolmis velatus)
WHITE MONJITA (Xolmis irupero)
BLACK-BACKED WATER-TYRANT (Fluvicola albiventer)
WHITE-HEADED MARSH TYRANT (Arundinicola leucocephala)
CATTLE TYRANT (Machetornis rixosa)
DULL-CAPPED ATTILA (Attila bolivianus)
RUFOUS CASIORNIS (Casiornis rufus)
SWAINSON'S FLYCATCHER (Myiarchus swainsoni)
SHORT-CRESTED FLYCATCHER (Myiarchus ferox)
BROWN-CRESTED FLYCATCHER (Myiarchus tyrannulus)
GREAT KISKADEE (Pitangus sulphuratus)
BOAT-BILLED FLYCATCHER (Megarynchus pitangua)
RUSTY-MARGINED FLYCATCHER (Myiozetetes cayanensis)
SOCIAL FLYCATCHER (Myiozetetes similis)
STREAKED FLYCATCHER (SOLITARIUS) (Myiodynastes maculatus solitarius)
TROPICAL KINGBIRD (Tyrannus melancholicus)
FORK-TAILED FLYCATCHER (Tyrannus savana)
Pipridae (Manakins)
SULPHUR-BELLIED TYRANT-MANAKIN (Neopelma sulphureiventer)
BAND-TAILED MANAKIN (Pipra fasciicauda)
Tityridae (Tityras and Allies)
GREEN-BACKED BECARD (Pachyramphus viridis)
Vireonidae (Vireos)
RUFOUS-BROWED PEPPERSHRIKE (Cyclarhis gujanensis)
Corvidae (Crows, Jays, and Magpies)
PURPLISH JAY (Cyanocorax cyanomelas)
PLUSH-CRESTED JAY (Cyanocorax chrysops)
Hirundinidae (Swallows)
TAWNY-HEADED SWALLOW (Alopochelidon fucata)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)
GRAY-BREASTED MARTIN (Progne chalybea)
BROWN-CHESTED MARTIN (Progne tapera)
WHITE-WINGED SWALLOW (Tachycineta albiventer)
WHITE-RUMPED SWALLOW (Tachycineta leucorrhoa)
Troglodytidae (Wrens)
THRUSH-LIKE WREN (Campylorhynchus turdinus unicolor)
FAWN-BREASTED WREN (Cantorchilus guarayanus)
Polioptilidae (Gnatcatchers)
MASKED GNATCATCHER (Polioptila dumicola berlepschi)
Donacobiidae (Donacobius)

Black-capped Donacobius was once thought to be a member of the wren family; looking at these birds with their throat sacs inflated, it's easy to accept that they belong in a family of their own. (Photo by guide Dan Lane)
BLACK-CAPPED DONACOBIUS (Donacobius atricapilla) Turdidae (Thrushes and Allies)
UNICOLORED THRUSH (Turdus haplochrous) [E]
CREAMY-BELLIED THRUSH (Turdus amaurochalinus)
Mimidae (Mockingbirds and Thrashers)
CHALK-BROWED MOCKINGBIRD (Mimus saturninus)
Motacillidae (Wagtails and Pipits)
YELLOWISH PIPIT (Anthus lutescens)
Parulidae (New World Warblers)
MASKED YELLOWTHROAT (Geothlypis aequinoctialis)
TROPICAL PARULA (Setophaga pitiayumi)
GOLDEN-CROWNED WARBLER (Basileuterus culicivorus)
Thraupidae (Tanagers and Allies)
CHESTNUT-VENTED CONEBILL (Conirostrum speciosum)
GUIRA TANAGER (Hemithraupis guira)
SILVER-BEAKED TANAGER (Ramphocelus carbo)
SAYACA TANAGER (Thraupis sayaca)
PALM TANAGER (Thraupis palmarum)
SWALLOW TANAGER (Tersina viridis)
GRAYISH SALTATOR (Saltator coerulescens)
Emberizidae (Buntings, Sparrows and Allies)
LONG-TAILED REED FINCH (Donacospiza albifrons)
BLUE-BLACK GRASSQUIT (Volatinia jacarina)
RUSTY-COLLARED SEEDEATER (Sporophila collaris)
WHITE-BELLIED SEEDEATER (Sporophila leucoptera bicolor)
SAFFRON FINCH (Sicalis flaveola)
WEDGE-TAILED GRASS-FINCH (Emberizoides herbicola)
GREAT PAMPA-FINCH (Embernagra platensis olivascens)
RED-CRESTED CARDINAL (Paroaria coronata)
RED-CAPPED CARDINAL (Paroaria gularis)
RED-CRESTED FINCH (Coryphospingus cucullatus)
YELLOW-BROWED SPARROW (Ammodramus aurifrons)
Icteridae (Troupials and Allies)
WHITE-BROWED BLACKBIRD (Sturnella superciliaris)
VELVET-FRONTED GRACKLE (Lampropsar tanagrinus boliviensis)
CHOPI BLACKBIRD (Gnorimopsar chopi)

Hard to miss this one: Orange-backed Troupial -- wow! (Photo by guide Dan Lane)
SCARLET-HEADED BLACKBIRD (Amblyramphus holosericeus) UNICOLORED BLACKBIRD (Agelasticus cyanopus)
BAY-WINGED COWBIRD (Agelaioides badius)
SHINY COWBIRD (Molothrus bonariensis)
GIANT COWBIRD (Molothrus oryzivorus)
VARIABLE ORIOLE (Icterus pyrrhopterus)
ORANGE-BACKED TROUPIAL (Icterus croconotus)
SOLITARY BLACK CACIQUE (Cacicus solitarius)
YELLOW-RUMPED CACIQUE (Cacicus cela)
CRESTED OROPENDOLA (Psarocolius decumanus)
Fringillidae (Siskins, Crossbills, and Allies)
PURPLE-THROATED EUPHONIA (Euphonia chlorotica)
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus)
FISHING BAT SP. (Noctilio albiventris)
BROWN CAPUCHIN (Cebus apella)
CAPYBARA (Hydrochaeris hydrochaeris)
AMAZON RIVER DOLPHIN (Inia geoffrensis boliviensis)
CRAB-EATING FOX (Cerdocyon thous)
SOUTH AMERICAN COATI (Nasua nasua)
Bat sp.
Tegu Lizard (Tupinambis sp.)
Spectacled Caiman (Caiman crocodilus)
Green Anaconda (Eunectes murinus)
Toilet Seat Frog (Trachycephalus typhonius)
Cane Toad (Bufo marinus)
'Leopard' Frog (Leptodactylus sp.)
Totals for the tour: 234 bird taxa and 6 mammal taxa