Enjoy our triplist below! For our tour
description, itinerary, past triplists, dates, fees, and more,
please visit our TOUR PAGE.
Click on the link at right to see this
triplist in printable PDF format with media only on
page 1.
The spiffy
Black-and-yellow Broadbill, though a fairly common Sundaland
specialty, turned out to be one of our favorites. (Photo by
tour participant Steve Rannels)
I've said it before, but I'm still thinking: what could be
more fun?! Take an enthusiastic group of birders to some of
the world's most impressive forests, full of exotic birds and
mammals, "flying" reptiles and amphibians, and a long list of
other incredible critters and plants, and we're sure to have a
great time. And so it was on this, our sixth, tour of some of
the best of Borneo. Dave and I thoroughly enjoyed it.
In our efforts to overcome jetlag, we all arrived early, and
we managed to get in a little extra birding before the tour
officially started. We had a morning to the N of KK, in open
country marshes and agricultural fields on the Tempusak Plain;
it produced a number of more widespread Asian species that we
don't see on the tour itself but that were clearly enjoyed by
this group, a number of whom were experiencing Asian birding
for the first time. (Since we all participated, I've included
those species in the list below.)
We began officially in the Crocker Range, where we saw a
number of highland endemics, from the usually tough
Whitehead's Broadbill and Mountain Serpent-Eagle to Mountain
and Bornean barbets, Bornean Treepie, Bornean Bulbul, Bornean
Leafbird, Black-sided Flowerpecker, and endearing flocks of
Chestnut-crested Yuhinas. We were off to a nice start.
At Gungung Kinabalu, Borneo's center of endemism, we had
wonderful luck with such specialties as the gorgeous
Whitehead's Trogon (a male before breakfast our very first
morning!), Fruit-hunter (including near an active nest),
Crimson-headed Partridge, Bare-headed Laughingthrush, Cave
Swiftlet, and such skulkers as Mountain Wren-Babbler, Bornean
Stubtail, and (for the Summit Trail group) Friendly
Bush-Warbler. We never heard even a note from a Whitehead's
Spiderhunter, and we failed to encounter the little-known
Everett's Thrush, but most of the rest of the endemics fell
into place, and we had a wonderful Rafflesia experience at
Poring and saw fabulous pitcher plants, including (for the
Mesilau group) the world's largest. Each morning we marveled
at the diversity of beautiful critters, especially moths, that
had accumulated outside Steve's door.
Our real introduction to tropical lowland forest was at
Sepilok, where we spent a morning in the RDC canopy towers,
enjoying a range of representative Asian species, from our
first hornbills and Black-and-yellow Broadbills to displaying
Violet Cuckoos, stunning White-bellied and Crimson-winged
woodpeckers, and fancy Raffles's and Chestnut-breasted
malkohas.
That afternoon we would continue to Gomantong Cave, where we
scoped out each of the echo-locating swiftlets on their nests
and then emerged from the cave to our first Orangutan! We
watched a long-winged Black Eagle sailing along the high
limestone cliffs and watched our first Red Leaf Monkeys
accomplishing some impressive jumps. We saw a calling pair of
White-crested Hornbills and watched large groups of
Bushy-crested Hornbills gather to roost sites as we waited for
the bats to emerge. Soon there were smoke-like clouds of
hundreds of Wrinkle-lipped Bats drifting away from the caves,
and shortly the crepuscular Bat Hawks appeared, diving through
the clouds and emerging with food for the night. We would
later see them perched and even hear their calls. It was a
lovely clear evening, and we managed to reach Sukau earlier
than usual.
We had sunny mornings, cloudy afternoons, and rainy evenings
for our two full days in the Kinabatangan. We birded by boat,
morning, late afternoon, and one evening, and we worked in
some time on the boardwalk immediately behind our lodge.
Highlights included the Bornean Ground-Cuckoo that we all
managed to see on our very first morning; electric light on
such beauties as Long-tailed Parakeets, Black-and-red
Broadbills, and Storm's Storks; the dramatic interactions
among three wing-lifting Great Slaty Woodpeckers; scope views
of Ruddy Kingfisher and Striped Wren-Babbler from our Sukau
boardwalk; a pair of tiny White-fronted Falconets perched up
beside the big Kinabatangan just before the heavens opened up;
incredible Proboscis Monkeys gathered along the riverbanks; a
Colugo with a young one that roosted during the day on a trunk
at our lodge and then sailed from one trunk to another by
night; and finding "our own" herd of Bornean Pygmy Elephants
at close range.
We had a wonderful morning of birding along the Gomantong
entrance road, seeing such wonders as Hooded and Black-headed
pittas, Red-bearded Bee-eater, and a terrific array of
babblers, before settling in to Borneo Rainforest Lodge, our
comfortable home for the next five days.
Foremost among our favorites at BRL were great views of the
monotypic Bornean Bristleheads on our first morning afield;
being able to watch at leisure a stunning male Blue-headed
Pitta repeatedly gathering dead-leaf material from the forest
floor and carrying it to a nest in the crotch of a tree 25
feet up!; and watching (and listening to!) the most bizarre of
the hornbills, the Helmeted. The canopy walkway offered
non-stop activity from time to time, with unique opportunities
to see canopy species eye-to-eye, and both the road and the
trails held ever-changing surprises, both by day and by night.
Among the many incredible birds, mammals, and other critters
we encountered at BRL, we'll each have our own additional
favorites. It was fun reliving the trip while annotating the
list that follows. Hopefully it'll serve to bring back great
memories for you guys as well.
It was a bit sobering to realize how many of the species we
saw are considered either Endangered, Vulnerable, or Near
Threatened by Birdlife International and the IUCN Red List.
We'll hope that our visit will contribute a bit toward their
longterm preservation.
In the annotations below, I've used the following
abbreviations:
RDC = Rainforest Discovery CentreBRL = Borneo Rainforest
Lodge
By a Sundaland specialty, I mean a species whose range is
generally restricted to the Thai-Malay peninsula, Borneo,
Sumatra, and Java‚ all of which were connected during the last
ice age. The eastern boundary of Sundaland is Wallace's Line,
a deep-water trench between Borneo and Sulawesi and between
Bali and Lombok, east of which is Wallacea/Australasia. Many
of the birds of Borneo, as marked with an S on the checklist,
are Sundaland specialties. Taxonomy follows the Clements
checklist with online updates. To access the Clements
checklist, go
tohttp://www.birds.cornell.edu/clementschecklist, where you
can download it and check for additional updates.
We thank Borneo Eco Tours, our great local guides (Adrian,
Haswan, Wangkong, and Paul), our captains and boatmen, and the
staffs of our wonderful lodgings throughout the tour. Special
thanks for cover photos to John Drummond (Fruit-hunter,
Black-sided Flowerpecker), Steve Rannels (Antheraea moth,
Black-and-yellow Broadbill), and Karen Walz (Orangutan). Dave
contributed the Storm's Stork, Streaky-breasted Spiderhunter,
and Blue-headed Pitta, and the rest were mine. Thanks, too, to
each of you guys for coming and for making the trip so much
fun.
--Rose Ann (& Dave)
For more information about this tour, including future
departures, visit our website at www.fieldguides.com.
And to see this same triplist online, go to https://fieldguides.com/triplists/bor12aLIST.pdf
and you will find the list in its entirety.
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for
individual species as appropriate: * = heard only, I =
introduced, E = endemic, N = nesting, a = austral migrant, b =
boreal migrant
Anatidae (Ducks, Geese, and Waterfowl)
LESSER WHISTLING-DUCK (Dendrocygna javanica)
Phasianidae (Pheasants, Grouse, and
Allies)
RED-BREASTED PARTRIDGE (Arborophila hyperythra) [E]
CHESTNUT-NECKLACED
PARTRIDGE (Arborophila
charltonii)
CRIMSON-HEADED
PARTRIDGE (Haematortyx
sanguiniceps) [E]
We found a very
friendly Friendly Bush-Warbler up the Summit Trail at
Kinabalu. (Photo by guide Dave Stejskal)
CRESTED FIREBACK (Lophura ignita nobilis)
GREAT ARGUS
(Argusianus argus)
Ciconiidae (Storks)
STORM'S STORK (Ciconia stormi)
LESSER ADJUTANT
(Leptoptilos javanicus)
Anhingidae (Anhingas)
ORIENTAL DARTER (Anhinga melanogaster)
Ardeidae (Herons, Egrets, and Bitterns)
YELLOW BITTERN (Ixobrychus sinensis)
CINNAMON BITTERN
(Ixobrychus cinnamomeus)
GRAY HERON
(Ardea cinerea)
GREAT-BILLED HERON
(Ardea sumatrana)
PURPLE HERON
(Ardea purpurea)
GREAT EGRET
(Ardea alba)
INTERMEDIATE EGRET
(Mesophoyx intermedia)
LITTLE EGRET
(Egretta garzetta)
PACIFIC REEF-HERON
(Egretta sacra)
CATTLE EGRET
(ASIAN) (Bubulcus ibis
coromandus) [b]
CHINESE POND-HERON
(Ardeola bacchus) [b]
JAVAN POND-HERON
(Ardeola speciosa)
[b]
STRIATED HERON
(Butorides striata)
BLACK-CROWNED
NIGHT-HERON (Nycticorax
nycticorax)
Accipitridae (Hawks, Eagles, and Kites)
JERDON'S BAZA (Aviceda jerdoni)
ORIENTAL
HONEY-BUZZARD (Pernis
ptilorhynchus)
BAT HAWK (Macheiramphus alcinus)
BLACK-SHOULDERED
KITE (Elanus caeruleus)
BRAHMINY KITE
(Haliastur indus)
WHITE-BELLIED
SEA-EAGLE (Haliaeetus
leucogaster)
LESSER FISH-EAGLE
(Ichthyophaga humilis)
MOUNTAIN
SERPENT-EAGLE (Spilornis
kinabaluensis) [E]
CRESTED
SERPENT-EAGLE (Spilornis
cheela)
CRESTED GOSHAWK
(Accipiter trivirgatus)
JAPANESE
SPARROWHAWK (Accipiter
gularis)
[b]
BLACK EAGLE
(Ictinaetus malayensis)
RUFOUS-BELLIED
EAGLE (Lophotriorchis
kienerii)
CHANGEABLE
HAWK-EAGLE (Nisaetus
cirrhatus) [N]
BLYTH'S HAWK-EAGLE
(Nisaetus alboniger)
Falconidae (Falcons and Caracaras)
WHITE-FRONTED FALCONET (Microhierax latifrons) [E]
PEREGRINE FALCON
(Falco peregrinus)
Rallidae (Rails, Gallinules, and Coots)
WHITE-BREASTED WATERHEN (Amaurornis phoenicurus)
WATERCOCK (Gallicrex cinerea) [b]
PURPLE SWAMPHEN
(BLACK-BACKED) (Porphyrio
porphyrio indicus)
EURASIAN MOORHEN
(Gallinula chloropus)
Charadriidae (Plovers and Lapwings)
PACIFIC GOLDEN-PLOVER (Pluvialis fulva) [b]
Recurvirostridae (Stilts and Avocets)
PIED STILT (Himantopus
leucocephalus) [a]
Scolopacidae (Sandpipers and Allies)
The canopy
walkway at BRL offers a totally different perspective on
canopy birding. (Photo by guide Rose Ann Rowlett)
COMMON SANDPIPER (Actitis hypoleucos) [b]
GREEN SANDPIPER
(Tringa ochropus) [b]
COMMON GREENSHANK
(Tringa nebularia)
MARSH SANDPIPER
(Tringa stagnatilis) [b]
WOOD SANDPIPER
(Tringa glareola) [b]
LONG-TOED STINT
(Calidris subminuta) [b]
PIN-TAILED SNIPE
(Gallinago stenura) [b]
Glareolidae (Pratincoles and Coursers)
ORIENTAL PRATINCOLE (Glareola maldivarum) [b]
Laridae (Gulls, Terns, and Skimmers)
GULL-BILLED TERN (Gelochelidon nilotica) [b]
WHITE-WINGED TERN
(Chlidonias leucopterus) [b]
WHISKERED TERN
(Chlidonias hybrida) [b]
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
SPOTTED DOVE
(Streptopelia chinensis)
RUDDY CUCKOO-DOVE
(Macropygia emiliana)
LITTLE CUCKOO-DOVE
(Macropygia ruficeps)
EMERALD DOVE
(Chalcophaps indica)
ZEBRA DOVE
(Geopelia striata) [I]
LITTLE
GREEN-PIGEON (Treron olax)
PINK-NECKED PIGEON
(Treron vernans)
LARGE GREEN-PIGEON
(Treron capellei)
GREEN
IMPERIAL-PIGEON (Ducula aenea)
MOUNTAIN
IMPERIAL-PIGEON (Ducula badia)
Psittacidae (Parrots)
BLUE-RUMPED PARROT (Psittinus cyanurus) [*]
LONG-TAILED
PARAKEET (Psittacula
longicauda)
BLUE-CROWNED
HANGING-PARROT (Loriculus
galgulus)
Cuculidae (Cuckoos)
LARGE HAWK-CUCKOO (DARK) (Hierococcyx sparverioides bocki) [*]
MOUSTACHED
HAWK-CUCKOO (Hierococcyx
vagans)
MALAYSIAN
HAWK-CUCKOO (Hierococcyx
fugax)
INDIAN CUCKOO
(Cuculus micropterus)
SUNDA CUCKOO
(Cuculus lepidus)
BANDED BAY CUCKOO
(Cacomantis sonneratii) [*]
PLAINTIVE CUCKOO
(Cacomantis merulinus)
LITTLE
BRONZE-CUCKOO (Chrysococcyx
minutillus)
VIOLET CUCKOO
(Chrysococcyx xanthorhynchus)
ASIAN
DRONGO-CUCKOO (Surniculus
lugubris)
ASIAN KOEL
(Eudynamys scolopaceus) [b]
BLACK-BELLIED
MALKOHA (Phaenicophaeus
diardi)
RAFFLES'S MALKOHA
(Phaenicophaeus chlorophaeus)
RED-BILLED MALKOHA
(Phaenicophaeus javanicus)
CHESTNUT-BREASTED
MALKOHA (Phaenicophaeus
curvirostris)
BORNEAN
GROUND-CUCKOO (Carpococcyx
radiatus) [E]
GREATER COUCAL
(Centropus sinensis)
LESSER COUCAL
(Centropus bengalensis)
Tytonidae (Barn-Owls)
ORIENTAL BAY-OWL (Phodilus badius) [*]
Strigidae (Owls)
MOUNTAIN SCOPS-OWL (Otus spilocephalus)
[*]
BUFFY FISH-OWL
(Ketupa ketupu)
BROWN WOOD-OWL
(Strix leptogrammica)
Apodidae (Swifts)
SILVER-RUMPED NEEDLETAIL (Rhaphidura leucopygialis)
BROWN-BACKED
NEEDLETAIL (Hirundapus
giganteus)
GLOSSY SWIFTLET
(Collocalia esculenta) [N]
CAVE SWIFTLET
(BORNEAN) (Collocalia linchi
dodgei) [N]
MOSSY-NEST
SWIFTLET (Aerodramus
salangana) [N]
BLACK-NEST
SWIFTLET (Aerodramus maximus)
EDIBLE-NEST
SWIFTLET (Aerodramus
fuciphagus)
The scarce
Red-throated Sunbird replaces Brown-throated Sunbird
inside primary forest at BRL. (Photo by tour participant
Steve Rannels)
HOUSE SWIFT (Apus nipalensis)
ASIAN PALM-SWIFT
(Cypsiurus balasiensis)
Hemiprocnidae (Treeswifts)
GRAY-RUMPED TREESWIFT (Hemiprocne longipennis)
WHISKERED
TREESWIFT (Hemiprocne comata)
Trogonidae (Trogons)
RED-NAPED TROGON (Harpactes kasumba)
DIARD'S TROGON
(Harpactes diardii)
WHITEHEAD'S TROGON
(Harpactes whiteheadi)
[E]
SCARLET-RUMPED
TROGON (Harpactes duvaucelii)
Alcedinidae (Kingfishers)
COMMON KINGFISHER (Alcedo atthis) [b]
BLUE-EARED
KINGFISHER (Alcedo meninting)
BLUE-BANDED
KINGFISHER (Alcedo euryzona)
[*]
RUFOUS-BACKED
KINGFISHER (Ceyx rufidorsa)
STORK-BILLED
KINGFISHER (Pelargopsis
capensis)
RUDDY KINGFISHER
(Halcyon coromanda)
COLLARED
KINGFISHER (Todiramphus
chloris)
RUFOUS-COLLARED
KINGFISHER (Actenoides
concretus) [*]
Meropidae (Bee-eaters)
RED-BEARDED BEE-EATER (Nyctyornis amictus)
BLUE-THROATED
BEE-EATER (Merops viridis)
[N]
Coraciidae (Rollers)
DOLLARBIRD (Eurystomus
orientalis)
Bucerotidae (Hornbills)
ORIENTAL PIED-HORNBILL (Anthracoceros albirostris)
BLACK HORNBILL
(Anthracoceros malayanus)
RHINOCEROS
HORNBILL (Buceros rhinoceros)
HELMETED HORNBILL
(Buceros vigil)
BUSHY-CRESTED
HORNBILL (Anorrhinus
galeritus)
WHITE-CROWNED
HORNBILL (Aceros comatus)
WRINKLED HORNBILL
(Aceros corrugatus)
WREATHED HORNBILL
(Aceros undulatus)
Megalaimidae (Asian Barbets)
BROWN BARBET (Calorhamphus fuliginosus)
GOLD-WHISKERED
BARBET (Megalaima chrysopogon)
RED-THROATED
BARBET (Megalaima
mystacophanos)
MOUNTAIN BARBET
(Megalaima monticola) [E]
YELLOW-CROWNED
BARBET (Megalaima henricii)
[*]
GOLDEN-NAPED
BARBET (Megalaima pulcherrima)
[E]
BLUE-EARED BARBET
(Megalaima australis) [N]
BORNEAN BARBET
(Megalaima eximia) [E]
Picidae (Woodpeckers)
RUFOUS PICULET (Sasia abnormis)
GRAY-CAPPED
WOODPECKER (Dendrocopos
canicapillus)
RUFOUS WOODPECKER
(Celeus brachyurus)
WHITE-BELLIED
WOODPECKER (Dryocopus
javensis)
BANDED WOODPECKER
(Picus miniaceus)
CRIMSON-WINGED
WOODPECKER (Picus puniceus)
CHECKER-THROATED
WOODPECKER (Picus mentalis)
MAROON WOODPECKER
(Blythipicus rubiginosus)
ORANGE-BACKED
WOODPECKER (Reinwardtipicus
validus)
BUFF-RUMPED
WOODPECKER (Meiglyptes
tristis)
BUFF-NECKED
WOODPECKER (Meiglyptes tukki)
GRAY-AND-BUFF
WOODPECKER (Hemicircus
concretus)
GREAT SLATY
WOODPECKER (Mulleripicus
pulverulentus)
Calyptomenidae (African and Green
Broadbills)
GREEN BROADBILL (Calyptomena viridis) [*]
WHITEHEAD'S
BROADBILL (Calyptomena
whiteheadi) [E]
Eurylaimidae (Asian and Grauer's
Broadbills)
BLACK-AND-RED BROADBILL (Cymbirhynchus macrorhynchos) [N]
LONG-TAILED
BROADBILL (Psarisomus
dalhousiae)
BANDED BROADBILL
(Eurylaimus javanicus) [N]
This male
Blue-headed Pitta was busy building a nest 25' above
ground in primary forest at BRL. (Photo by tour
participant John Drummond)
BLACK-AND-YELLOW BROADBILL (Eurylaimus ochromalus) [N]
DUSKY BROADBILL
(Corydon sumatranus)
Pittidae (Pittas)
GIANT PITTA (Pitta caerulea)
BANDED PITTA
(BORNEAN) (Pitta guajana
schwaneri)
BLUE-HEADED PITTA
(Pitta baudii) [EN]
BLACK-HEADED PITTA
(Pitta ussheri) [E]
HOODED PITTA
(Pitta sordida)
Acanthizidae (Thornbills and Allies)
GOLDEN-BELLIED GERYGONE (Gerygone sulphurea) [*]
Prionopidae (Helmetshrikes and Allies)
LARGE WOODSHRIKE (Tephrodornis gularis)
RUFOUS-WINGED
PHILENTOMA (Philentoma
pyrhoptera)
MAROON-BREASTED
PHILENTOMA (Philentoma velata)
Artamidae (Woodswallows)
WHITE-BREASTED WOODSWALLOW (Artamus leucorynchus)
Pityriaseidae (Bristlehead)
BORNEAN BRISTLEHEAD (Pityriasis gymnocephala)
[E]
Aegithinidae (Ioras)
COMMON IORA (Aegithina tiphia)
GREEN IORA
(Aegithina viridissima)
Campephagidae (Cuckoo-shrikes)
SUNDA CUCKOO-SHRIKE (Coracina larvata)
LESSER
CUCKOO-SHRIKE (Coracina
fimbriata)
PIED TRILLER
(Lalage nigra)
FIERY MINIVET
(Pericrocotus igneus)
SCARLET MINIVET
(Pericrocotus flammeus)
GRAY-CHINNED
MINIVET (Pericrocotus solaris)
BAR-WINGED
FLYCATCHER-SHRIKE (Hemipus
picatus)
BLACK-WINGED
FLYCATCHER-SHRIKE (Hemipus
hirundinaceus)
[N]
Pachycephalidae (Whistlers and Allies)
BORNEAN WHISTLER (Pachycephala hypoxantha) [E]
Laniidae (Shrikes)
BROWN SHRIKE (Lanius cristatus) [b]
LONG-TAILED SHRIKE
(Lanius schach)
Vireonidae (Vireos)
BLYTH'S SHRIKE-BABBLER (BLYTH'S)
(Pteruthius aeralatus robinsoni)
Oriolidae (Old World Orioles)
DARK-THROATED ORIOLE (Oriolus xanthonotus)
BLACK-AND-CRIMSON
ORIOLE (Oriolus cruentus)
Dicruridae (Drongos)
ASHY DRONGO (Dicrurus leucophaeus)
CROW-BILLED DRONGO
(Dicrurus annectans)
[b]
BRONZED DRONGO
(Dicrurus aeneus)
HAIR-CRESTED
DRONGO (Dicrurus hottentottus)
GREATER
RACKET-TAILED DRONGO (Dicrurus
paradiseus brachyphorus)
Rhipiduridae (Fantails)
WHITE-THROATED FANTAIL (Rhipidura albicollis)
PIED FANTAIL
(Rhipidura javanica)
SPOTTED FANTAIL
(Rhipidura perlata)
Monarchidae (Monarch Flycatchers)
BLACK-NAPED MONARCH (Hypothymis azurea)
ASIAN
PARADISE-FLYCATCHER (Terpsiphone
paradisi)
Corvidae (Crows, Jays, and Magpies)
CRESTED JAY (Platylophus galericulatus coronatus)
BLACK MAGPIE
(Platysmurus leucopterus aterrimus)
SHORT-TAILED
MAGPIE (Cissa thalassina)
BORNEAN TREEPIE
(Dendrocitta cinerascens) [E]
SLENDER-BILLED
CROW (Corvus enca)
Hirundinidae (Swallows)
BARN SWALLOW (Hirundo rustica)
[b]
PACIFIC SWALLOW
(Hirundo tahitica) [N]
STRIATED SWALLOW
(Cecropis striolata) [b]
Stenostiridae (Fairy Flycatchers)
GRAY-HEADED CANARY-FLYCATCHER (Culicicapa ceylonensis)
Sittidae (Nuthatches)
VELVET-FRONTED NUTHATCH (Sitta frontalis)
Pycnonotidae (Bulbuls)
PUFF-BACKED BULBUL (Pycnonotus eutilotus) [*]
BLACK-HEADED
BULBUL (Pycnonotus atriceps)
BORNEAN BULBUL
(Pycnonotus montis) [E]
SCALY-BREASTED
BULBUL (Pycnonotus squamatus)
FLAVESCENT BULBUL
(Pycnonotus flavescens leucops)
YELLOW-VENTED
BULBUL (Pycnonotus goiavier)
OLIVE-WINGED
BULBUL (Pycnonotus plumosus)
RED-EYED BULBUL
(Pycnonotus brunneus)
SPECTACLED BULBUL
(Pycnonotus erythropthalmos)
HAIRY-BACKED
BULBUL (Tricholestes criniger)
FINSCH'S BULBUL
(Alophoixus finschii)
OCHRACEOUS BULBUL
(Alophoixus ochraceus)
GRAY-CHEEKED
BULBUL (Alophoixus bres)
YELLOW-BELLIED
BULBUL (Alophoixus
phaeocephalus)
BUFF-VENTED BULBUL
(Iole olivacea)
ASHY BULBUL
(CINEREOUS) (Hemixos flavala
connectens)
STREAKED BULBUL
(Ixos malaccensis)
Cettiidae (Bush-Warblers and Allies)
BORNEAN STUBTAIL (Urosphena whiteheadi) [E]
YELLOW-BELLIED
WARBLER (Abroscopus
superciliaris)
MOUNTAIN
TAILORBIRD (Phyllergates
cucullatus)
SUNDA BUSH-WARBLER
(Cettia vulcania)
Phylloscopidae (Leaf-Warblers)
ARCTIC WARBLER (Phylloscopus borealis) [b]
MOUNTAIN WARBLER
(Phylloscopus trivirgatus)
The handsome Red
Leaf Monkey is one of 10 sympatric primates in the
Kinabatangan. (Photo by guide Dave Stejskal)
YELLOW-BREASTED WARBLER (Seicercus montis)
Acrocephalidae (Reed-Warblers and Allies)
ORIENTAL REED-WARBLER (Acrocephalus orientalis) [b]
Locustellidae (Grassbirds and Allies)
FRIENDLY BUSH-WARBLER (Bradypterus accentor) [E]
STRIATED GRASSBIRD
(Megalurus palustris)
Cisticolidae (Cisticolas and Allies)
DARK-NECKED TAILORBIRD (Orthotomus atrogularis)
RUFOUS-TAILED
TAILORBIRD (Orthotomus
sericeus)
ASHY TAILORBIRD
(Orthotomus ruficeps)
YELLOW-BELLIED
PRINIA (Prinia flaviventris)
Zosteropidae (Yuhinas, White-eyes, and
Allies)
CHESTNUT-CRESTED YUHINA (Yuhina everetti) [E]
PYGMY WHITE-EYE
(Oculocincta squamifrons)
[E]
MOUNTAIN BLACK-EYE
(Chlorocharis emiliae)
[E]
BLACK-CAPPED
WHITE-EYE (Zosterops
atricapilla)
Pellorneidae (Fulvettas and Ground
Babblers)
BROWN FULVETTA (Alcippe brunneicauda)
BLACK-THROATED
WREN-BABBLER (Turdinus
atrigularis) [E]
HORSFIELD'S
BABBLER (Malacocincla
sepiaria)
SHORT-TAILED
BABBLER (Malacocincla
malaccensis)
STRIPED
WREN-BABBLER (Kenopia striata)
TEMMINCK'S BABBLER
(Pellorneum pyrrogenys)
BLACK-CAPPED
BABBLER (Pellorneum
capistratum)
WHITE-CHESTED
BABBLER (Trichastoma
rostratum)
FERRUGINOUS
BABBLER (Trichastoma bicolor)
MOUSTACHED BABBLER
(Malacopteron magnirostre)
SOOTY-CAPPED
BABBLER (Malacopteron affine)
SCALY-CROWNED
BABBLER (Malacopteron
cinereum)
RUFOUS-CROWNED
BABBLER (Malacopteron magnum)
MOUNTAIN
WREN-BABBLER (Napothera
crassa) [E]
BORNEAN
WREN-BABBLER (Ptilocichla
leucogrammica) [E*]
Leiothrichidae (Laughingthrushes)
SUNDA LAUGHINGTHRUSH (Garrulax palliatus)
BARE-HEADED
LAUGHINGTHRUSH (Garrulax
calvus) [E]
CHESTNUT-CAPPED
LAUGHINGTHRUSH (CHESTNUT-HOODED) (Garrulax mitratus treacheri)
Timaliidae (Babblers)
RUFOUS-FRONTED BABBLER (Stachyridopsis rufifrons)
BOLD-STRIPED
TIT-BABBLER (Macronous
bornensis) [E]
FLUFFY-BACKED
TIT-BABBLER (Macronous
ptilosus)
GRAY-THROATED
BABBLER (Stachyris nigriceps)
GRAY-HEADED
BABBLER (Stachyris
poliocephala)
BLACK-THROATED
BABBLER (Stachyris
nigricollis)
CHESTNUT-RUMPED
BABBLER (Stachyris maculata)
CHESTNUT-WINGED
BABBLER (Stachyris
erythroptera)
CHESTNUT-BACKED
SCIMITAR-BABBLER (Pomatorhinus
montanus)
Irenidae (Fairy-bluebirds)
ASIAN FAIRY-BLUEBIRD (Irena puella)
Muscicapidae (Old World Flycatchers)
DARK-SIDED FLYCATCHER (Muscicapa sibirica) [b]
GRAY-STREAKED
FLYCATCHER (Muscicapa
griseisticta) [b]
ASIAN BROWN
FLYCATCHER (Muscicapa
dauurica) [b]
ORIENTAL
MAGPIE-ROBIN (Copsychus
saularis)
RUFOUS-TAILED
SHAMA (Copsychus pyrropygus)
WHITE-RUMPED SHAMA
(WHITE-CROWNED) (Copsychus
malabaricus stricklandii)
PALE
BLUE-FLYCATCHER (Cyornis
unicolor) [*]
LONG-BILLED
BLUE-FLYCATCHER (Cyornis
caerulatus)
MALAYSIAN
BLUE-FLYCATCHER (Cyornis
turcosus)
BORNEAN
BLUE-FLYCATCHER (Cyornis
superbus) [E]
GRAY-CHESTED
JUNGLE-FLYCATCHER (Cyornis
umbratilis)
PYGMY
BLUE-FLYCATCHER (Muscicapella
hodgsoni)
BLUE-AND-WHITE
FLYCATCHER (Cyanoptila
cyanomelana) [b]
INDIGO FLYCATCHER
(Eumyias indigo)
EYEBROWED
JUNGLE-FLYCATCHER (Vauriella
gularis) [E]
WHITE-BROWED
SHORTWING (Brachypteryx
montana)
BORNEAN
WHISTLING-THRUSH (Myophonus
borneensis) [E]
The largest
flower in Borneo, Rafflesia keithii has no leaves or
stems; it is totally parasitic on an underground vine.
(Photo by tour participant Karen Walz)
WHITE-CROWNED FORKTAIL (Enicurus leschenaulti frontalis)
WHITE-CROWNED
FORKTAIL (Enicurus
leschenaulti borneensis)
CHESTNUT-NAPED
FORKTAIL (Enicurus
ruficapillus)
RUFOUS-CHESTED
FLYCATCHER (Ficedula
dumetoria)
LITTLE PIED
FLYCATCHER (Ficedula
westermanni)
MUGIMAKI
FLYCATCHER (Ficedula mugimaki)
[b]
SNOWY-BROWED
FLYCATCHER (Ficedula
hyperythra)
Turdidae (Thrushes and Allies)
EYEBROWED THRUSH (Turdus obscurus) [b]
FRUIT-HUNTER
(Chlamydochaera jefferyi) [EN]
Sturnidae (Starlings)
ASIAN GLOSSY STARLING (Aplonis panayensis)
COMMON HILL MYNA
(Gracula religiosa)
JAVAN MYNA
(Acridotheres javanicus) [I]
Chloropseidae (Leafbirds)
GREATER GREEN LEAFBIRD (Chloropsis sonnerati)
LESSER GREEN
LEAFBIRD (Chloropsis
cyanopogon)
BORNEAN LEAFBIRD
(Chloropsis kinabaluensis) [E]
Dicaeidae (Flowerpeckers)
YELLOW-BREASTED FLOWERPECKER (Prionochilus maculatus)
YELLOW-RUMPED
FLOWERPECKER (Prionochilus
xanthopygius) [E]
THICK-BILLED
FLOWERPECKER (Dicaeum agile)
BROWN-BACKED
FLOWERPECKER (Dicaeum
everetti)
YELLOW-VENTED
FLOWERPECKER (Dicaeum
chrysorrheum)
ORANGE-BELLIED
FLOWERPECKER (Dicaeum
trigonostigma)
BLACK-SIDED
FLOWERPECKER (Dicaeum
monticolum) [E]
SCARLET-BACKED
FLOWERPECKER (Dicaeum
cruentatum)
Nectariniidae (Sunbirds and Spiderhunters)
RUBY-CHEEKED SUNBIRD (Chalcoparia singalensis)
PLAIN SUNBIRD
(Anthreptes simplex)
PLAIN-THROATED
SUNBIRD (Anthreptes
malacensis)
RED-THROATED
SUNBIRD (Anthreptes
rhodolaemus)
PURPLE-NAPED
SUNBIRD (Hypogramma
hypogrammicum)
VAN HASSELT'S
SUNBIRD (Leptocoma brasiliana)
OLIVE-BACKED
SUNBIRD (Cinnyris jugularis)
EASTERN CRIMSON
SUNBIRD (Aethopyga siparaja)
TEMMINCK'S SUNBIRD
(Aethopyga temminckii)
LITTLE
SPIDERHUNTER (Arachnothera
longirostra)
THICK-BILLED
SPIDERHUNTER (Arachnothera
crassirostris)
LONG-BILLED
SPIDERHUNTER (Arachnothera
robusta)
SPECTACLED
SPIDERHUNTER (Arachnothera
flavigaster)
YELLOW-EARED
SPIDERHUNTER (Arachnothera
chrysogenys)
STREAKY-BREASTED
SPIDERHUNTER (Arachnothera
affinis everetti)
Motacillidae (Wagtails and Pipits)
EASTERN YELLOW WAGTAIL (Motacilla tschutschensis simillima)
[b]
WHITE WAGTAIL
(OCULARIS) (Motacilla alba
ocularis) [b]
ORIENTAL PIPIT
(Anthus rufulus malayensis)
Passeridae (Old World Sparrows)
EURASIAN TREE SPARROW (Passer montanus)
[I]
Estrildidae (Waxbills and Allies)
DUSKY MUNIA (Lonchura fuscans) [E]
NUTMEG MANNIKIN
(Lonchura punctulata)
WHITE-BELLIED
MUNIA (Lonchura leucogastra)
[N]
CHESTNUT MUNIA
(Lonchura atricapilla)
COLUGO (Cynocephalus variegatus)
WRINKLE-LIPPED BAT
(Tadarida [Chaerephon] plicata)
COMMON TREESHREW
(Tupaia glis)
MOUNTAIN TREESHREW
(Tupaia montana) [E]
This Mountain
Treeshrew could be contributing critical nitrogen to the
largest pitcher plant on Earth. (Photo by tour participant
Steve Rannels)
SLOW LORIS (Nycticebus
cougang)
CRAB-EATING
MACAQUE (Macaca fascigularis)
PIGTAIL MACAQUE
(Macaca nemestrina)
SILVERED LEAF
MONKEY (Presbytis cristata)
RED LEAF MONKEY
(Presbytis rubicunda) [E]
PROBOSCIS MONKEY
(Nasalis larvatus) [E]
GRAY GIBBON
(Hylobates muelleri) [E]
ORANGUTAN (Pongo pygmaeus) [E]
GIANT SQUIRREL SP.
(Ratufa affinis)
PREVOST'S SQUIRREL
(Callosciurus prevostii)
PLANTAIN SQUIRREL
(Callosciurus notatus)
BORNEAN
BLACK-BANDED SQUIRREL (Callosciurus
orestes) [E]
JENTINK'S SQUIRREL
(Sundasciurus jentincki) [E]
BORNEAN MOUNTAIN
GROUND-SQUIRREL (Dremomys
everetti) [E]
PLAIN PYGMY
SQUIRREL (Exilisciurus exilis)
[E]
RED GIANT FLYING
SQUIRREL (Petaurista
petaurista)
THOMAS'S FLYING
SQUIRREL (Aeromys thomasi)
[E]
COMMON PORCUPINE
(Hystrix brachyura)
SMALL-CLAWED
(INDIAN SMOOTH) OTTER (Aonyx
cinerea)
MALAY CIVET
(Viverra tangalunga)
MASKED PALM CIVET
(Paguma larvata)
BINTURONG (Arctictis binturong)
BANDED PALM CIVET
(Hemigalus derbyanus)
BORNEAN PYGMY
ELEPHANT (Elephas maximus
borneensis)
BEARDED PIG
(Sus barbatus)
GREATER MOUSE DEER
(Tragulus napu)
LESSER MOUSE DEER
(Tragulus javanicus)
MUNTJAC (BARKING
DEER) (Muntiacus muntjak)
SAMBAR (Cervus unicolor)
One of the joys of birding in Borneo is all the delightful
distractions that come in varied and exotic forms from other
parts of the animal kingdom. Our triplist database contains
only birds and mammals, but we had a long list of additional
write-ins, including a few plant species of exceptional
interest. To mention a very few, emphasizing those that we
were able to identify:
Among the plants:
--Rafflesia keithii - We traveled to Poring to see the flower
of this fascinating Bornean endemic, the largest of its genus
in Borneo. Adrian explained how scientists and the Kinabalu
Park staff work with locals to enhance the environmental
conditions favorable to this rare parasitic plant, namely by
making its host vine available for distribution on nearby
private lands. We saw several buds and a fabulous blossom on
its second day open, as well as several older, decaying
flowers.
--Carnivorous Nepenthes pitcher plants - The pitcher plant
center of Borneo (and the world!) is the upper-elevation Mt.
Kinabalu. This was the first year we've offered an alternative
trip to Mesilau, where we took a steep (but short) trail to 4
different species of pitcher plants: N. burbigeae (see my
cover photo), N. fusca (a small one), N. tentaculata (the same
one that was seen by the group birding the Summit Trail); and
N. rajah, a.k.a. Rajah Brooke's Pitcher Plant, the largest
pitcher plant in the world, its pitcher holding up to 3.5
liters of liquid! Recent research has revealed the
"evolutionary incentive" for high-elevation pitcher plants to
have evolved bigger and bigger pitchers: It's to serve as a
toilet for treeshrews! Treeshrews, which are common at these
high elevations (where insects become increasinly scarce) are
attracted by the sweet secretion on the underside of the
pitcher's lid. To reach it, they climb onto the pitcher's
sturdy rim, which fits them perfectly, and sip away, meanwhile
defecating into the pitcher to mark their feeding
territory--thus supplying all the nitrogen critical to these
normally carnivorous plants. See the following link for a full
explanation, with photos:
http://news.bbc.co.uk/earth/hi/earth_news/newsid_8552000/8552157.stm
Among the other vertebrates of particular interest were:
--that huge Water Monitor (Varanus salvator), in the crotch
of a tree along the Menanggul River
--the giant Tokay Geckos (Gekko gecko) that were usually
under the covered section of the boardwalk between the rooms
and the dining area at Sukau Rainforest Lodge; these were the
ones we heard throughout the lowlands, uttering the loud
"gec-ko" or "to-kay" that slows down; it was for this species
that the genus was named.
--the cool Kuhl's Parachute Gecko (Ptychozoon kuhli) that
some of us saw on the trunk of a tree at the Danum bridge;
those frilly extensions on its sides and all of its appendages
help it sail from tree to tree by night, when it's active.
--Blue-eyed Angle-headed Lizard (Gonocephalus liogaster) -
This is what I think the crested agamid lizard was that we saw
by night at BRL. Its nuchal and dorsal crests were continuous
and tall, and it showed a blue eye with yellow eyering. It's
another Sundaland specialty, found only in peninsular
Malaysia, Sumatra, and Borneo.
--small "flying lizards" of the genus Draco - The bright
green one with the pointed orange display pouch was probably
D. cornutus. They were seen at Sukau and BRL.
--Mangrove (or Yellow-banded) Cat Snake (Boiga dendrophila),
the shiny black snake with the brioght-yellow bands that was
so tightly coiled in a tree overhanging the Menanggul that we
could hardly see its head! Their diet consists of birds and
their eggs, amphibians, lizards, other snakes, mouse deer (!),
and treeshrews.
--Red-sided Keelback Water Snake (Xenochrophis
trianguligerus) This was what Paul called a "Red-sided Tree
Snake" when we saw and photographed it at the BRL "frog pond."
It's a mildly venomous member of the Colubridae that feeds
almost exclusively on frogs near bodies of water.
--File-eared Tree Frog (Polypedates otilophus), the richly
colored frogs also at the BRL frog pond. They occur only in
Sumatra and Borneo, ordinarily well above the ground. But they
congregate over water to breed and lay their fertilized eggs
into a foam mass that will harden on the outside while
remaining liquid on the inside, providing the tadpoles a
suitable environment until the mass falls into the water
below.
--Harlequin Flying Tree Frog (Rhacophorus pardalis). Also at
the BRL frog pond. Little did we realize at the time that this
little frog spends most of its time in the canopy, only coming
to the ground to breed in suitable streams and pools. It too
is a foam nest builder, but it's one of the "flying" frogs,
with extensive membranes between its toes to help it glide
from branch to branch--and down to pools on the forest floor.
It occurs in peninsular Malaysia, Sumatra, Borneo, and the
Philippines.
Among the many invertebrates were some that simply must be
remembered:
--the common Wood Nymph (Idea stolli) or Tree Nymph
butterflies that floated tissue paper-like throughout the
lowlands
--the striking black-and-yellow Common Birdwings (Trioides
helena) and many other superb butterflies [see the following
website to help ID some of your photos:
http://www.learnaboutbutterflies.com/Malaysia%20thumbs.htm]
--the many gorgeous moths attracted to Steve's black light at
Kinabalu, including the big beauty on the cover (Antheraea
sp.; thanks for the photo, Steve); and the big Lyssa moth
(Lyssa menoetcus) that was photographed along the boardwalk to
our rooms at BRL. We'll try to post the names of the ones
we've ID'd in a slideshow, to be compiled eventually.
--the long-snouted Chinese Lantern Bug (Pyrops sp.) that
Diana found at Poring (and I think some were seen at BRL as
well?). These are sap suckers, which excrete excess sugars as
droplets that certain ants have specialized to collect!
--those wonderful black-and-orange Trilobite Beetles
(Duliticola paradoxa; in rotten logs at Kinabalu) that retain
their larval form even when sexually mature
--that unbelievable praying mantis that was so elaborately
trimmed to resemble a dead stick with leafy extensions; all
the big cicadas, katydids, and other amazing insects in the
lowland rainforest
--the hordes of cockroaches and scuterigid centipedes that
decorated the walls of Gomantong Cave
--those long, dark millipedes with bright orange legs; and
big, short, fat Pill Millipedes (Glomeris connexa) that curl
into a balllike a pillbug, both fairly common at BRL
--some wonderful spiders, including lots of "wolf spiders"
with eyeshine and that big arboreal tarantula that Paul
pointed out on our final night drive
--and did I mention the leeches? Especially notable were
those yellow-striped Tiger Leeches (Haemadipsa picta) at BRL,
yet another Bornean specialty. They informed our fashion and
left several of us with a "battled scar" to show off at home.
Like Kinabalu itself, that's just the "tip of the batholith."
There are LOTS of fabulous critters out there, and we
encountered a wonderful sampling of them.
Totals for the tour: 328 bird taxa and 33 mammal taxa