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When it comes to tropical color, Costa Rica's many bright birds -- including this stunning male Red-legged Honeycreeper -- can't be beat. Photo by participant Paul Bisson.
Another year, another great trip to Costa Rica. You might think that after 30+ tours to this country, I'd be getting a little tired of it, but that is definitely not the case. Costa Rica is where my love of Neotropical birds was kindled more than 20 years ago, and I still love it. And this year's awesome tour has just reinforced that passion I have for this country and its birds.
We kicked things off in grand style at Virgen del Socorro on our first morning, picking up eye level views of a gorgeous pair of Barred Hawks circling over the road, then finding a fig tree laden with fruit, as well as howler monkeys and a great variety of birds: Bay-headed, Emerald, and Speckled tanagers, Scarlet-thighed Dacnis, Red-headed Barbet, Pale-vented Thrush, Golden-browed Chlorophonia, Crested Guan, and more. Retreating to La Paz Waterfall Gardens for lunch, we were enthralled by the numbers of cool hummingbirds at the feeders, not to mention the Sooty-faced Finches under our table, and Prong-billed Barbet, Spangle-cheeked Tanager, and Slaty-backed Nightingale-Thrush in the surrounding forest. It was a memorable first day in the field.
Things heated up (literally) in the Caribbean lowlands, and our time at La Selva was a whirlwind of wonderful birds: Great Curassows and a Great Tinamou strolling along the forest floor, a juvenile Gray-headed Kite gulping down a juicy green caterpillar, a White-fronted Nunbird calling noisily above the cabins, a bunch of stunning woodpecker species, including floppy-crested Chestnut-colored, and the huge Pale-billed, the latter pounding out his double-knock drum in full view of the group, swarms of parrots, including at least half a dozen gaudy Great Green Macaws, a trio of Purple-throated Fruitcrows, throats shimmering brightly in the sunlight, a tiny Black-capped Pygmy-Tyrant sitting quietly in the canopy, and others too numerous to mention here.
Rancho Naturalista was our next destination, but there were some good finds on our way there, too. A viewpoint over the endless forested slopes at Braulio Carrillo offered up scope views of a King Vulture and an Ornate Hawk-Eagle, while a White Hawk sat calmly over our heads, watching us as intently as we were watching it. The park trails held a large swirling flock of Tawny-crested and Carmiol's tanagers, and Black-headed Nightingale-Thrush, the local Lattice-tailed Trogon, and a pair of minuscule Red-fronted Parrotlets delighted those that saw them. Further down the road, a roosting Great Potoo was a big hit as well. Rancho, meanwhile gave us the usual fantastic hummingbirds (Snowcap, Crowned Woodnymph, Green Thorntail), the moth cloth (White-breasted Wood-wren, Red-throated Ant-Tanager, the scarce Tawny-chested Flycatcher, etc.) and the trails (lekking White-crowned Manakins, nesting Black-headed Tody-Flycatchers, and a pair of hungry Bicolored Hawks sharing an avian meal). A little further afield, a pair of spectacular Sunbitterns feeding two fuzzy youngsters in a nest was another area highlight.
Moving up into the highlands, we had a marvelous evening foray near Orosi, with easy Mottled Owls, and slightly more difficult Bare-shanked Screech-owls, plus a bonus Gray Fox to show for our efforts. In Tapanti the next day, we enjoyed the likes of a perched Green-fronted Lancebill, a somewhat stunned (though ultimately fine) Spotted Barbtail in the hand, showy Crimson-collared Tanagers and Black-thighed Grosbeak, and the rare sight of a Sharpbill working on its nest, only the second known nest of this species in the country! Higher still in the Savegre Valley, most of the montane specialties came through for us. We had an unforgettable encounter with a shimmering male Resplendent Quetzal as the indisputable highlight, but sandwiched around that sighting were equally wonderful encounters with such charismatic birds as Spotted Wood-Quail, Scaled Antpitta, Buffy Tuftedcheek, Streak-breasted Treehunter, Long-tailed Silky-Flycatcher, Wrenthrush, Collared Redstart, Yellow-thighed Finch, and plenty more.
Moving down to the Pacific coast and Carara National Park, we encountered a whole new avifauna, and new species came at an incredible rate. Gorgeous Scarlet Macaws, exquisite Turquoise-browed Motmots, beautiful Baird's Trogons, and flashy Fiery-billed Aracaris were among the more showy species to cross our paths, but it was a bunch of less colorful birds that stole the show here. That amazing army ant swarm at Carara was one of the best I've ever seen, and was easily one of the high points of this trip. Having birds like Black-faced Antthrush, Dusky, Chestnut-backed, and Bicolored antbirds, Black-hooded Antshrike, Buff-throated Foliage-gleaner, Rufous-and-white Wren, Gray-headed Tanager, and more strolling across and perching on the path was unbelievable! What an awesome experience! We finished up the birding with a morning in the dry tropical forest, complete with White-throated Magpie-Jay, Laughing Falcon, and Lesser Ground-Cuckoo, to name but a few, and a couple of days in the famous Monteverde area. Though we were short on targets by this time, we came away with things like Three-wattled Bellbird, Long-tailed Manakin, Brown-billed Scythebill, and Black-breasted Wood-Quail to round out our trip lists.
In closing, I just want to thank all of you for choosing to join me on this tour, and for being such a fun and compatible group of travelers. I love it when my job doesn't feel too much like work, and that's all down to you folks making it so easy on me. Thanks, too, to my co-leaders and friends, Ernesto and Vernon, for all their help in making this trip a great success, and to our able office staff, especially our tour manager, Caroline, for ensuring everything ran smoothly for us from the moment we hit the ground. I look forward to seeing you all on another tour sometime soon. In the meantime, get out and enjoy the spring migration. That's where I'm heading right now!
-- Jay
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant

Tinamous tend to be shy and retiring, so finding this Great Tinamou right out in the open was a real treat. Photo by participant Paul Bisson.
Tinamidae (Tinamous)
GREAT TINAMOU (Tinamus major)
Anatidae (Ducks, Geese, and Waterfowl)
BLACK-BELLIED WHISTLING-DUCK (Dendrocygna autumnalis)
MUSCOVY DUCK (Cairina moschata)
BLUE-WINGED TEAL (Anas discors) [b]
RING-NECKED DUCK (Aythya collaris) [b]
LESSER SCAUP (Aythya affinis) [b]
Cracidae (Guans, Chachalacas, and Curassows)
GRAY-HEADED CHACHALACA (Ortalis cinereiceps)
CRESTED GUAN (Penelope purpurascens)
BLACK GUAN (Chamaepetes unicolor)
GREAT CURASSOW (Crax rubra)
Odontophoridae (New World Quail)
BUFFY-CROWNED WOOD-PARTRIDGE (Dendrortyx leucophrys) [*]
BLACK-BREASTED WOOD-QUAIL (Odontophorus leucolaemus)
SPOTTED WOOD-QUAIL (Odontophorus guttatus)
Podicipedidae (Grebes)
LEAST GREBE (Tachybaptus dominicus)
Ciconiidae (Storks)
WOOD STORK (Mycteria americana)

Bare-faced Tiger-Herons were relatively common in the Pacific lowlands. Photo by participant Reg David.
Fregatidae (Frigatebirds)
MAGNIFICENT FRIGATEBIRD (Fregata magnificens)
Phalacrocoracidae (Cormorants and Shags)
NEOTROPIC CORMORANT (Phalacrocorax brasilianus)
Anhingidae (Anhingas)
ANHINGA (Anhinga anhinga)
Pelecanidae (Pelicans)
BROWN PELICAN (Pelecanus occidentalis)
Ardeidae (Herons, Egrets, and Bitterns)
LEAST BITTERN (Ixobrychus exilis)
FASCIATED TIGER-HERON (Tigrisoma fasciatum)
BARE-THROATED TIGER-HERON (Tigrisoma mexicanum)
GREAT BLUE HERON (Ardea herodias) [b]
GREAT EGRET (Ardea alba)
SNOWY EGRET (Egretta thula)
LITTLE BLUE HERON (Egretta caerulea)
TRICOLORED HERON (Egretta tricolor)
CATTLE EGRET (Bubulcus ibis)
GREEN HERON (Butorides virescens)
BLACK-CROWNED NIGHT-HERON (Nycticorax nycticorax)
YELLOW-CROWNED NIGHT-HERON (Nyctanassa violacea)
BOAT-BILLED HERON (Cochlearius cochlearius) [N]
Threskiornithidae (Ibises and Spoonbills)
WHITE IBIS (Eudocimus albus)

The gang heads out. Adventure awaits! Photo by participant Greg Vassilopoulos.
GREEN IBIS (Mesembrinibis cayennensis)
ROSEATE SPOONBILL (Platalea ajaja)
Cathartidae (New World Vultures)
BLACK VULTURE (Coragyps atratus)
TURKEY VULTURE (Cathartes aura)
KING VULTURE (Sarcoramphus papa)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus) [b]
Accipitridae (Hawks, Eagles, and Kites)
WHITE-TAILED KITE (Elanus leucurus)
GRAY-HEADED KITE (Leptodon cayanensis)
SWALLOW-TAILED KITE (Elanoides forficatus)
ORNATE HAWK-EAGLE (Spizaetus ornatus)
SNAIL KITE (Rostrhamus sociabilis)
DOUBLE-TOOTHED KITE (Harpagus bidentatus)
SHARP-SHINNED HAWK (Accipiter striatus) [b]
COOPER'S HAWK (Accipiter cooperii) [b]
BICOLORED HAWK (Accipiter bicolor) [N]
CRANE HAWK (Geranospiza caerulescens)

Gray-necked Wood-Rails tend to be a bit more confiding than most members of their family. Photo by participant Reg David.
COMMON BLACK HAWK (MANGROVE) (Buteogallus anthracinus subtilis) [N]
BARRED HAWK (Morphnarchus princeps)
ROADSIDE HAWK (Rupornis magnirostris)
WHITE HAWK (Pseudastur albicollis)
GRAY HAWK (Buteo plagiatus)
BROAD-WINGED HAWK (Buteo platypterus) [b]
SHORT-TAILED HAWK (Buteo brachyurus)
SWAINSON'S HAWK (Buteo swainsoni) [b]
ZONE-TAILED HAWK (Buteo albonotatus)
RED-TAILED HAWK (Buteo jamaicensis)
Eurypygidae (Sunbittern)
SUNBITTERN (Eurypyga helias) [N]
Rallidae (Rails, Gallinules, and Coots)
WHITE-THROATED CRAKE (Laterallus albigularis) [*]
GRAY-NECKED WOOD-RAIL (Aramides cajaneus)
PURPLE GALLINULE (Porphyrio martinicus)

Finding a Sunbittern on its nest (complete with two fluffy youngsters) was one of the highlights of the trip. Photo by participant Reg David.
Aramidae (Limpkin)
LIMPKIN (Aramus guarauna)
Burhinidae (Thick-knees)
DOUBLE-STRIPED THICK-KNEE (Burhinus bistriatus)
Charadriidae (Plovers and Lapwings)
BLACK-BELLIED PLOVER (Pluvialis squatarola) [b]
SOUTHERN LAPWING (Vanellus chilensis)
WILSON'S PLOVER (Charadrius wilsonia)
SEMIPALMATED PLOVER (Charadrius semipalmatus) [b]
KILLDEER (Charadrius vociferus) [b]
Jacanidae (Jacanas)
NORTHERN JACANA (Jacana spinosa)
Scolopacidae (Sandpipers and Allies)
SPOTTED SANDPIPER (Actitis macularius) [b]
WILLET (Tringa semipalmata) [b]
WHIMBREL (Numenius phaeopus) [b]
RUDDY TURNSTONE (Arenaria interpres) [b]
SANDERLING (Calidris alba) [b]
LEAST SANDPIPER (Calidris minutilla) [b]
SEMIPALMATED SANDPIPER (Calidris pusilla) [b]
Laridae (Gulls, Terns, and Skimmers)
LAUGHING GULL (Leucophaeus atricilla)
ROYAL TERN (Thalasseus maximus)
SANDWICH TERN (Thalasseus sandvicensis)

Our new nightbirding spot near Orosi proved very productive, with a Mottled Owl popping in to check things out when we squeaked for a passing fox. Photo by participant Reg David.
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
PALE-VENTED PIGEON (Patagioenas cayennensis)
RED-BILLED PIGEON (Patagioenas flavirostris)
BAND-TAILED PIGEON (Patagioenas fasciata)
RUDDY PIGEON (Patagioenas subvinacea)
SHORT-BILLED PIGEON (Patagioenas nigrirostris)
INCA DOVE (Columbina inca)
COMMON GROUND-DOVE (Columbina passerina)
RUDDY GROUND-DOVE (Columbina talpacoti)
WHITE-TIPPED DOVE (Leptotila verreauxi)
GRAY-CHESTED DOVE (Leptotila cassinii)
BUFF-FRONTED QUAIL-DOVE (Zentrygon costaricensis)
WHITE-WINGED DOVE (Zenaida asiatica)
Cuculidae (Cuckoos)
SQUIRREL CUCKOO (Piaya cayana)
LESSER GROUND-CUCKOO (Morococcyx erythropygus)
GROOVE-BILLED ANI (Crotophaga sulcirostris)
Tytonidae (Barn-Owls)
BARN OWL (Tyto alba)
Strigidae (Owls)
TROPICAL SCREECH-OWL (Megascops choliba)
VERMICULATED SCREECH-OWL (VERMICULATED) (Megascops guatemalae vermiculatus)
BARE-SHANKED SCREECH-OWL (Megascops clarkii)
CRESTED OWL (Lophostrix cristata) [*]
SPECTACLED OWL (Pulsatrix perspicillata)
COSTA RICAN PYGMY-OWL (Glaucidium costaricanum) [*]
FERRUGINOUS PYGMY-OWL (Glaucidium brasilianum)
MOTTLED OWL (Ciccaba virgata)
BLACK-AND-WHITE OWL (Ciccaba nigrolineata)
STRIPED OWL (Pseudoscops clamator)

Lesser Ground-Cuckoos can be tough to find if they're not singing; fortunately, this one was serenading the neighborhood from a shrubby mound right near the road. Photo by participant Reg David.
Caprimulgidae (Nightjars and Allies)
LESSER NIGHTHAWK (Chordeiles acutipennis)
SHORT-TAILED NIGHTHAWK (Lurocalis semitorquatus)
COMMON PAURAQUE (Nyctidromus albicollis)
DUSKY NIGHTJAR (Antrostomus saturatus)
Nyctibiidae (Potoos)
GREAT POTOO (Nyctibius grandis)
Apodidae (Swifts)
WHITE-COLLARED SWIFT (Streptoprocne zonaris)
VAUX'S SWIFT (Chaetura vauxi)
COSTA RICAN SWIFT (Chaetura fumosa)
GRAY-RUMPED SWIFT (Chaetura cinereiventris)
LESSER SWALLOW-TAILED SWIFT (Panyptila cayennensis)
Trochilidae (Hummingbirds)
WHITE-NECKED JACOBIN (Florisuga mellivora)
BAND-TAILED BARBTHROAT (Threnetes ruckeri)
GREEN HERMIT (Phaethornis guy)
LONG-BILLED HERMIT (Phaethornis longirostris)
STRIPE-THROATED HERMIT (Phaethornis striigularis)
GREEN-FRONTED LANCEBILL (Doryfera ludovicae) [N]
BROWN VIOLETEAR (Colibri delphinae)
GREEN VIOLETEAR (Colibri thalassinus)
PURPLE-CROWNED FAIRY (Heliothryx barroti)
GREEN-BREASTED MANGO (Anthracothorax prevostii)
GREEN THORNTAIL (Discosura conversii)

A Great Potoo on its dayroost demonstrates its top-notch imitation of a tree stump. Photo by participant Paul Bisson.
GREEN-CROWNED BRILLIANT (Heliodoxa jacula)
MAGNIFICENT HUMMINGBIRD (Eugenes fulgens)
PLAIN-CAPPED STARTHROAT (Heliomaster constantii)
FIERY-THROATED HUMMINGBIRD (Panterpe insignis)
WHITE-BELLIED MOUNTAIN-GEM (Lampornis hemileucus)
PURPLE-THROATED MOUNTAIN-GEM (Lampornis calolaemus)
WHITE-THROATED MOUNTAIN-GEM (GRAY-TAILED) (Lampornis castaneoventris cinereicauda)
MAGENTA-THROATED WOODSTAR (Calliphlox bryantae)
RUBY-THROATED HUMMINGBIRD (Archilochus colubris) [b]
VOLCANO HUMMINGBIRD (Selasphorus flammula)
SCINTILLANT HUMMINGBIRD (Selasphorus scintilla)
VIOLET-HEADED HUMMINGBIRD (Klais guimeti)
SCALY-BREASTED HUMMINGBIRD (Phaeochroa cuvierii)
VIOLET SABREWING (Campylopterus hemileucurus)
BRONZE-TAILED PLUMELETEER (Chalybura urochrysia)
CROWNED WOODNYMPH (Thalurania colombica)
STRIPE-TAILED HUMMINGBIRD (Eupherusa eximia)
BLACK-BELLIED HUMMINGBIRD (Eupherusa nigriventris)
WHITE-TAILED EMERALD (Elvira chionura)
COPPERY-HEADED EMERALD (Elvira cupreiceps) [E]
SNOWCAP (Microchera albocoronata)
BLUE-CHESTED HUMMINGBIRD (Amazilia amabilis)
CHARMING HUMMINGBIRD (Amazilia decora)
MANGROVE HUMMINGBIRD (Amazilia boucardi) [E]
STEELY-VENTED HUMMINGBIRD (Amazilia saucerottei)
RUFOUS-TAILED HUMMINGBIRD (Amazilia tzacatl) [N]

A female Purple-throated Mountaingem doesn't even bend the edge of the leaf she's sitting on. Photo by participant Reg David.
CINNAMON HUMMINGBIRD (Amazilia rutila)
Trogonidae (Trogons)
RESPLENDENT QUETZAL (Pharomachrus mocinno)
LATTICE-TAILED TROGON (Trogon clathratus)
SLATY-TAILED TROGON (Trogon massena)
BLACK-HEADED TROGON (Trogon melanocephalus)
BAIRD'S TROGON (Trogon bairdii)
GARTERED TROGON (Trogon caligatus)
BLACK-THROATED TROGON (Trogon rufus)
ORANGE-BELLIED TROGON (Trogon aurantiiventris)
COLLARED TROGON (Trogon collaris)
Momotidae (Motmots)
BLUE-CROWNED MOTMOT (LESSON'S) (Momotus coeruliceps lessonii)
RUFOUS MOTMOT (Baryphthengus martii)
BROAD-BILLED MOTMOT (Electron platyrhynchum)
TURQUOISE-BROWED MOTMOT (Eumomota superciliosa)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)
BELTED KINGFISHER (Megaceryle alcyon) [b]
AMAZON KINGFISHER (Chloroceryle amazona)
GREEN KINGFISHER (Chloroceryle americana)
AMERICAN PYGMY KINGFISHER (Chloroceryle aenea)

The dapper White-necked Jacobin is typically one of the feistier birds around the feeders. Photo by participant Paul Bisson.
Bucconidae (Puffbirds)
WHITE-NECKED PUFFBIRD (Notharchus hyperrhynchus)
WHITE-WHISKERED PUFFBIRD (Malacoptila panamensis)
WHITE-FRONTED NUNBIRD (Monasa morphoeus)
Galbulidae (Jacamars)
RUFOUS-TAILED JACAMAR (Galbula ruficauda)
Capitonidae (New World Barbets)
RED-HEADED BARBET (Eubucco bourcierii)
Semnornithidae (Toucan-Barbets)
PRONG-BILLED BARBET (Semnornis frantzii)
Ramphastidae (Toucans)
EMERALD TOUCANET (BLUE-THROATED) (Aulacorhynchus prasinus caeruleogularis)
COLLARED ARACARI (Pteroglossus torquatus)
FIERY-BILLED ARACARI (Pteroglossus frantzii)
YELLOW-THROATED TOUCAN (CHESTNUT-MANDIBLED) (Ramphastos ambiguus swainsonii)
KEEL-BILLED TOUCAN (Ramphastos sulfuratus)
Picidae (Woodpeckers)
OLIVACEOUS PICULET (Picumnus olivaceus)
ACORN WOODPECKER (Melanerpes formicivorus)
GOLDEN-NAPED WOODPECKER (Melanerpes chrysauchen)
BLACK-CHEEKED WOODPECKER (Melanerpes pucherani)
RED-CROWNED WOODPECKER (Melanerpes rubricapillus)
HOFFMANN'S WOODPECKER (Melanerpes hoffmannii)
SMOKY-BROWN WOODPECKER (Picoides fumigatus) [*]
HAIRY WOODPECKER (Picoides villosus)
RUFOUS-WINGED WOODPECKER (Piculus simplex)
GOLDEN-OLIVE WOODPECKER (Colaptes rubiginosus)
CINNAMON WOODPECKER (Celeus loricatus)
CHESTNUT-COLORED WOODPECKER (Celeus castaneus)
LINEATED WOODPECKER (Dryocopus lineatus) [N]
PALE-BILLED WOODPECKER (Campephilus guatemalensis)
Falconidae (Falcons and Caracaras)
CRESTED CARACARA (Caracara cheriway)

When you get a close look, it's easy to see how the Green Violetear got its name! Photo by participant Paul Bisson.
YELLOW-HEADED CARACARA (Milvago chimachima)
LAUGHING FALCON (Herpetotheres cachinnans)
BAT FALCON (Falco rufigularis)
PEREGRINE FALCON (Falco peregrinus) [b]
Psittacidae (New World and African Parrots)
RED-FRONTED PARROTLET (Touit costaricensis)
BARRED PARAKEET (Bolborhynchus lineola)
ORANGE-CHINNED PARAKEET (Brotogeris jugularis)
BROWN-HOODED PARROT (Pyrilia haematotis)
WHITE-CROWNED PARROT (Pionus senilis)
RED-LORED PARROT (Amazona autumnalis)
YELLOW-NAPED PARROT (Amazona auropalliata)
WHITE-FRONTED PARROT (Amazona albifrons)
MEALY PARROT (Amazona farinosa)
SULPHUR-WINGED PARAKEET (Pyrrhura hoffmanni)
OLIVE-THROATED PARAKEET (AZTEC) (Eupsittula nana astec)
ORANGE-FRONTED PARAKEET (Eupsittula canicularis)

It's clearly hard work being a Scintillant Hummingbird! Photo by participant Paul Bisson.
GREAT GREEN MACAW (Ara ambiguus)
SCARLET MACAW (Ara macao) [N]
CRIMSON-FRONTED PARAKEET (Psittacara finschi)
Thamnophilidae (Typical Antbirds)
FASCIATED ANTSHRIKE (Cymbilaimus lineatus)
BARRED ANTSHRIKE (Thamnophilus doliatus)
BLACK-HOODED ANTSHRIKE (Thamnophilus bridgesi)
RUSSET ANTSHRIKE (Thamnistes anabatinus)
PLAIN ANTVIREO (Dysithamnus mentalis)
STREAK-CROWNED ANTVIREO (Dysithamnus striaticeps) [*]
CHECKER-THROATED ANTWREN (Epinecrophylla fulviventris)
SLATY ANTWREN (Myrmotherula schisticolor)
DOT-WINGED ANTWREN (Microrhopias quixensis)
DUSKY ANTBIRD (Cercomacroides tyrannina)
CHESTNUT-BACKED ANTBIRD (Myrmeciza exsul)
DULL-MANTLED ANTBIRD (Myrmeciza laemosticta) [*]
ZELEDON'S ANTBIRD (Myrmeciza zeledoni)
BICOLORED ANTBIRD (Gymnopithys bicolor bicolor)
Grallariidae (Antpittas)
SCALED ANTPITTA (Grallaria guatimalensis)

Its easy to see why Resplendent Quetzal won the "bird of the trip" honors, hands down. What a stunner! Photo by participant Greg Vassilopoulos.
Rhinocryptidae (Tapaculos)
SILVERY-FRONTED TAPACULO (Scytalopus argentifrons)
Formicariidae (Antthrushes)
BLACK-FACED ANTTHRUSH (Formicarius analis)
Furnariidae (Ovenbirds and Woodcreepers)
TAWNY-THROATED LEAFTOSSER (Sclerurus mexicanus) [*]
OLIVACEOUS WOODCREEPER (Sittasomus griseicapillus)
TAWNY-WINGED WOODCREEPER (Dendrocincla anabatina)
PLAIN-BROWN WOODCREEPER (Dendrocincla fuliginosa)
WEDGE-BILLED WOODCREEPER (Glyphorynchus spirurus)
NORTHERN BARRED-WOODCREEPER (Dendrocolaptes sanctithomae)
COCOA WOODCREEPER (Xiphorhynchus susurrans)
BLACK-STRIPED WOODCREEPER (Xiphorhynchus lachrymosus) [*]
SPOTTED WOODCREEPER (Xiphorhynchus erythropygius)
BROWN-BILLED SCYTHEBILL (Campylorhamphus pusillus)
STREAK-HEADED WOODCREEPER (Lepidocolaptes souleyetii)
SPOT-CROWNED WOODCREEPER (Lepidocolaptes affinis)
PLAIN XENOPS (Xenops minutus)
BUFFY TUFTEDCHEEK (Pseudocolaptes lawrencii)
LINEATED FOLIAGE-GLEANER (Syndactyla subalaris)
STREAK-BREASTED TREEHUNTER (Thripadectes rufobrunneus)
BUFF-THROATED FOLIAGE-GLEANER (HYPOPHAEUS) (Automolus ochrolaemus hypophaeus)
BUFF-THROATED FOLIAGE-GLEANER (EXSERTUS) (Automolus ochrolaemus exsertus)
STRIPED WOODHAUNTER (Automolus subulatus) [*]
SPOTTED BARBTAIL (Premnoplex brunnescens)
RUDDY TREERUNNER (Margarornis rubiginosus)
RED-FACED SPINETAIL (Cranioleuca erythrops)

An Emerald Toucanet catching the sun by the feeders at La Cinchona shows nicely why this subspecies (caeruleogularis) is called "Blue-throated Toucanet." Photo by participant Reg David.
Tyrannidae (Tyrant Flycatchers)
NORTHERN BEARDLESS-TYRANNULET (Camptostoma imberbe)
SOUTHERN BEARDLESS-TYRANNULET (Camptostoma obsoletum)
GREENISH ELAENIA (Myiopagis viridicata)
YELLOW-BELLIED ELAENIA (Elaenia flavogaster)
MOUNTAIN ELAENIA (Elaenia frantzii)
TORRENT TYRANNULET (Serpophaga cinerea)
OLIVE-STRIPED FLYCATCHER (Mionectes olivaceus)
OCHRE-BELLIED FLYCATCHER (Mionectes oleagineus)
SLATY-CAPPED FLYCATCHER (Leptopogon superciliaris)
PALTRY TYRANNULET (Zimmerius vilissimus)
NORTHERN SCRUB-FLYCATCHER (Sublegatus arenarum arenarum)
BLACK-CAPPED PYGMY-TYRANT (Myiornis atricapillus)
SCALE-CRESTED PYGMY-TYRANT (Lophotriccus pileatus)
NORTHERN BENTBILL (Oncostoma cinereigulare)
SLATE-HEADED TODY-FLYCATCHER (Poecilotriccus sylvia)

Acorn Woodpeckers are plentiful -- and noisily obvious! -- around our hotel in the Savegre valley. Photo by participant Paul Bisson.
COMMON TODY-FLYCATCHER (Todirostrum cinereum)
BLACK-HEADED TODY-FLYCATCHER (Todirostrum nigriceps)
EYE-RINGED FLATBILL (Rhynchocyclus brevirostris)
YELLOW-OLIVE FLYCATCHER (Tolmomyias sulphurescens)
STUB-TAILED SPADEBILL (Platyrinchus cancrominus) [*]
GOLDEN-CROWNED SPADEBILL (Platyrinchus coronatus)
ROYAL FLYCATCHER (NORTHERN) (Onychorhynchus coronatus mexicanus) [N]
RUDDY-TAILED FLYCATCHER (Terenotriccus erythrurus)
SULPHUR-RUMPED FLYCATCHER (Myiobius sulphureipygius aureatus)
TAWNY-CHESTED FLYCATCHER (Aphanotriccus capitalis)
TUFTED FLYCATCHER (Mitrephanes phaeocercus)
OLIVE-SIDED FLYCATCHER (Contopus cooperi) [b]
DARK PEWEE (Contopus lugubris)
OCHRACEOUS PEWEE (Contopus ochraceus)
WESTERN WOOD-PEWEE (Contopus sordidulus) [b]
EASTERN WOOD-PEWEE (Contopus virens) [b]
TROPICAL PEWEE (Contopus cinereus)
YELLOW-BELLIED FLYCATCHER (Empidonax flaviventris) [b]
YELLOWISH FLYCATCHER (Empidonax flavescens)
BLACK-CAPPED FLYCATCHER (Empidonax atriceps)
BLACK PHOEBE (Sayornis nigricans)
LONG-TAILED TYRANT (Colonia colonus)
BRIGHT-RUMPED ATTILA (Attila spadiceus)
RUFOUS MOURNER (Rhytipterna holerythra)
DUSKY-CAPPED FLYCATCHER (Myiarchus tuberculifer)
PANAMA FLYCATCHER (Myiarchus panamensis)
NUTTING'S FLYCATCHER (Myiarchus nuttingi)
GREAT CRESTED FLYCATCHER (Myiarchus crinitus) [b]
BROWN-CRESTED FLYCATCHER (Myiarchus tyrannulus)

Scarlet Macaws are now hard to miss in the Carara region. Photo by participant Paul Bisson.
GREAT KISKADEE (Pitangus sulphuratus) [N]
BOAT-BILLED FLYCATCHER (Megarynchus pitangua)
SOCIAL FLYCATCHER (Myiozetetes similis)
GRAY-CAPPED FLYCATCHER (Myiozetetes granadensis) [N]
WHITE-RINGED FLYCATCHER (Conopias albovittatus)
GOLDEN-BELLIED FLYCATCHER (Myiodynastes hemichrysus)
STREAKED FLYCATCHER (Myiodynastes maculatus)
SULPHUR-BELLIED FLYCATCHER (Myiodynastes luteiventris) [a]
PIRATIC FLYCATCHER (Legatus leucophaius) [a]
TROPICAL KINGBIRD (Tyrannus melancholicus)
SCISSOR-TAILED FLYCATCHER (Tyrannus forficatus) [b]
Oxyruncidae (Sharpbill)
SHARPBILL (Oxyruncus cristatus) [N]
Cotingidae (Cotingas)
PURPLE-THROATED FRUITCROW (Querula purpurata)
RUFOUS PIHA (Lipaugus unirufus)
THREE-WATTLED BELLBIRD (Procnias tricarunculatus)
Pipridae (Manakins)
LONG-TAILED MANAKIN (Chiroxiphia linearis)
WHITE-RUFFED MANAKIN (Corapipo altera)
BLUE-CROWNED MANAKIN (Lepidothrix coronata)
WHITE-COLLARED MANAKIN (Manacus candei)
ORANGE-COLLARED MANAKIN (Manacus aurantiacus)
WHITE-CROWNED MANAKIN (Dixiphia pipra)
RED-CAPPED MANAKIN (Ceratopipra mentalis)
Tityridae (Tityras and Allies)
MASKED TITYRA (Tityra semifasciata)

An American Crocodile lurks on a river bank. Photo by participant Reg David.
BARRED BECARD (Pachyramphus versicolor)
CINNAMON BECARD (Pachyramphus cinnamomeus)
ROSE-THROATED BECARD (Pachyramphus aglaiae)
Vireonidae (Vireos, Shrike-Babblers, and Erpornis)
MANGROVE VIREO (Vireo pallens)
YELLOW-THROATED VIREO (Vireo flavifrons) [b]
YELLOW-WINGED VIREO (Vireo carmioli)
BROWN-CAPPED VIREO (Vireo leucophrys)
PHILADELPHIA VIREO (Vireo philadelphicus) [b]
YELLOW-GREEN VIREO (Vireo flavoviridis) [a]
TAWNY-CROWNED GREENLET (Tunchiornis ochraceiceps)
LESSER GREENLET (Pachysylvia decurtata)
RUFOUS-BROWED PEPPERSHRIKE (Cyclarhis gujanensis) [*]
Corvidae (Crows, Jays, and Magpies)
WHITE-THROATED MAGPIE-JAY (Calocitta formosa)
BROWN JAY (Psilorhinus morio)
Hirundinidae (Swallows)
BLUE-AND-WHITE SWALLOW (Pygochelidon cyanoleuca)

Yeah, the exposure is a little off, and the picture is a little grainy. But it's a picture of a Sharpbill on a nest, and there are only two known nesting records for the entire country! Photo by participant Reg David.
NORTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx serripennis)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)
MANGROVE SWALLOW (Tachycineta albilinea) [N]
BARN SWALLOW (Hirundo rustica) [b]
Troglodytidae (Wrens)
NIGHTINGALE WREN (Microcerculus philomela) [*]
SCALY-BREASTED WREN (WHISTLING) (Microcerculus marginatus luscinia) [*]
HOUSE WREN (Troglodytes aedon)
OCHRACEOUS WREN (Troglodytes ochraceus) [N]
TIMBERLINE WREN (Thryorchilus browni)
BAND-BACKED WREN (Campylorhynchus zonatus)
RUFOUS-NAPED WREN (Campylorhynchus rufinucha)
BLACK-BELLIED WREN (Pheugopedius fasciatoventris)
RUFOUS-BREASTED WREN (Pheugopedius rutilus)
BLACK-THROATED WREN (Pheugopedius atrogularis)
BANDED WREN (Thryophilus pleurostictus)
RUFOUS-AND-WHITE WREN (Thryophilus rufalbus)
STRIPE-BREASTED WREN (Cantorchilus thoracicus)

We had many great looks at Passerini's Tanagers on the Caribbean slope. Photo by participant Paul Bisson.
PLAIN WREN (Cantorchilus modestus)
RIVERSIDE WREN (Cantorchilus semibadius)
BAY WREN (Cantorchilus nigricapillus)
WHITE-BREASTED WOOD-WREN (Henicorhina leucosticta)
GRAY-BREASTED WOOD-WREN (Henicorhina leucophrys)
Polioptilidae (Gnatcatchers)
LONG-BILLED GNATWREN (Ramphocaenus melanurus)
WHITE-LORED GNATCATCHER (Polioptila albiloris)
TROPICAL GNATCATCHER (Polioptila plumbea)
Cinclidae (Dippers)
AMERICAN DIPPER (Cinclus mexicanus)
Turdidae (Thrushes and Allies)
BLACK-FACED SOLITAIRE (Myadestes melanops) [N]
BLACK-BILLED NIGHTINGALE-THRUSH (Catharus gracilirostris)
ORANGE-BILLED NIGHTINGALE-THRUSH (Catharus aurantiirostris)
SLATY-BACKED NIGHTINGALE-THRUSH (Catharus fuscater)

Green Honeycreepers are found on both slopes in Costa Rica, but we found most of ours this year (including this female) on the Caribbean side. Photo by participant Reg David.
RUDDY-CAPPED NIGHTINGALE-THRUSH (Catharus frantzii)
BLACK-HEADED NIGHTINGALE-THRUSH (Catharus mexicanus)
SWAINSON'S THRUSH (Catharus ustulatus) [b]
WOOD THRUSH (Hylocichla mustelina) [b]
SOOTY THRUSH (Turdus nigrescens)
MOUNTAIN THRUSH (Turdus plebejus)
PALE-VENTED THRUSH (Turdus obsoletus)
CLAY-COLORED THRUSH (Turdus grayi)
WHITE-THROATED THRUSH (Turdus assimilis)
Mimidae (Mockingbirds and Thrashers)
TROPICAL MOCKINGBIRD (Mimus gilvus)
Ptiliogonatidae (Silky-flycatchers)
BLACK-AND-YELLOW SILKY-FLYCATCHER (Phainoptila melanoxantha)
LONG-TAILED SILKY-FLYCATCHER (Ptiliogonys caudatus)
Parulidae (New World Warblers)
OVENBIRD (Seiurus aurocapilla) [b]
WORM-EATING WARBLER (Helmitheros vermivorum) [b]
LOUISIANA WATERTHRUSH (Parkesia motacilla) [b]
NORTHERN WATERTHRUSH (Parkesia noveboracensis) [b]

The Blue-gray Tanager was among the handful of species we saw every day of the tour. Photo by participant Paul Bisson.
GOLDEN-WINGED WARBLER (Vermivora chrysoptera) [b]
BLACK-AND-WHITE WARBLER (Mniotilta varia) [b]
PROTHONOTARY WARBLER (Protonotaria citrea) [b]
FLAME-THROATED WARBLER (Oreothlypis gutturalis)
TENNESSEE WARBLER (Oreothlypis peregrina) [b]
GRAY-CROWNED YELLOWTHROAT (Geothlypis poliocephala)
KENTUCKY WARBLER (Geothlypis formosa) [b]
OLIVE-CROWNED YELLOWTHROAT (Geothlypis semiflava)
TROPICAL PARULA (Setophaga pitiayumi)
BLACKBURNIAN WARBLER (Setophaga fusca) [b]
YELLOW WARBLER (Setophaga petechia) [b]
YELLOW WARBLER (MANGROVE) (Setophaga petechia erithachorides)
CHESTNUT-SIDED WARBLER (Setophaga pensylvanica) [b]
BLACK-THROATED GREEN WARBLER (Setophaga virens) [b]
RUFOUS-CAPPED WARBLER (Basileuterus rufifrons)
BLACK-CHEEKED WARBLER (Basileuterus melanogenys)
GOLDEN-CROWNED WARBLER (Basileuterus culicivorus)
THREE-STRIPED WARBLER (Basileuterus tristriatus)

A male Slaty Flowerpiercer pauses between nectar raids, showing the sharply pointed beak which allows him to cut his way into the flower. This little one is a thief -- stealing nectar without pollinating the plant. Photo by participant Reg David.
BUFF-RUMPED WARBLER (Myiothlypis fulvicauda)
WILSON'S WARBLER (Cardellina pusilla) [b]
SLATE-THROATED REDSTART (Myioborus miniatus)
COLLARED REDSTART (Myioborus torquatus)
WRENTHRUSH (Zeledonia coronata)
Thraupidae (Tanagers and Allies)
GRAY-HEADED TANAGER (Eucometis penicillata)
WHITE-SHOULDERED TANAGER (Tachyphonus luctuosus)
TAWNY-CRESTED TANAGER (Tachyphonus delatrii)
WHITE-LINED TANAGER (Tachyphonus rufus)
CRIMSON-COLLARED TANAGER (Ramphocelus sanguinolentus)
PASSERINI'S TANAGER (Ramphocelus passerinii)
CHERRIE'S TANAGER (Ramphocelus costaricensis)
BLUE-GRAY TANAGER (Thraupis episcopus)
PALM TANAGER (Thraupis palmarum)
GOLDEN-HOODED TANAGER (Tangara larvata)
SPECKLED TANAGER (Tangara guttata)
SPANGLE-CHEEKED TANAGER (Tangara dowii)

Chestnut-headed Oropendola is the less common of Costa Rica's two oropendola species. Photo by participant Paul Bisson.
PLAIN-COLORED TANAGER (Tangara inornata)
BAY-HEADED TANAGER (Tangara gyrola)
EMERALD TANAGER (Tangara florida)
SILVER-THROATED TANAGER (Tangara icterocephala) [N]
SCARLET-THIGHED DACNIS (Dacnis venusta)
BLUE DACNIS (Dacnis cayana)
RED-LEGGED HONEYCREEPER (Cyanerpes cyaneus)
GREEN HONEYCREEPER (Chlorophanes spiza)
BLACK-AND-YELLOW TANAGER (Chrysothlypis chrysomelas) [*]
SLATY FLOWERPIERCER (Diglossa plumbea)
BLUE-BLACK GRASSQUIT (Volatinia jacarina)
THICK-BILLED SEED-FINCH (Sporophila funerea)
NICARAGUAN SEED-FINCH (Sporophila nuttingi)
VARIABLE SEEDEATER (Sporophila corvina)
WHITE-COLLARED SEEDEATER (Sporophila torqueola)
BANANAQUIT (Coereba flaveola) [N]
YELLOW-FACED GRASSQUIT (Tiaris olivaceus)
DUSKY-FACED TANAGER (Mitrospingus cassinii)
BUFF-THROATED SALTATOR (Saltator maximus)
BLACK-HEADED SALTATOR (Saltator atriceps)
GRAYISH SALTATOR (Saltator coerulescens)
Emberizidae (Buntings and New World Sparrows)
SOOTY-CAPPED CHLOROSPINGUS (Chlorospingus pileatus)
COMMON CHLOROSPINGUS (Chlorospingus flavopectus)
STRIPE-HEADED SPARROW (Peucaea ruficauda)
OLIVE SPARROW (Arremonops rufivirgatus)
ORANGE-BILLED SPARROW (Arremon aurantiirostris)
CHESTNUT-CAPPED BRUSHFINCH (Arremon brunneinucha)
SOOTY-FACED FINCH (Arremon crassirostris)
VOLCANO JUNCO (Junco vulcani)
RUFOUS-COLLARED SPARROW (Zonotrichia capensis)

We found our Volcano Junco, a high-elevation specialist typically found above treeline, on Cerro de la Muerte. Photo by participant Paul Bisson.
LARGE-FOOTED FINCH (Pezopetes capitalis)
WHITE-EARED GROUND-SPARROW (Melozone leucotis)
YELLOW-THIGHED FINCH (Pselliophorus tibialis)
WHITE-NAPED BRUSHFINCH (YELLOW-THROATED) (Atlapetes albinucha gutturalis)
Cardinalidae (Cardinals and Allies)
HEPATIC TANAGER (Piranga flava)
SUMMER TANAGER (Piranga rubra) [b]
FLAME-COLORED TANAGER (Piranga bidentata)
RED-THROATED ANT-TANAGER (Habia fuscicauda)
CARMIOL'S TANAGER (Chlorothraupis carmioli)
BLACK-FACED GROSBEAK (Caryothraustes poliogaster)
BLACK-THIGHED GROSBEAK (Pheucticus tibialis)
ROSE-BREASTED GROSBEAK (Pheucticus ludovicianus) [b]
BLUE-BLACK GROSBEAK (Cyanocompsa cyanoides)
BLUE GROSBEAK (Passerina caerulea)
PAINTED BUNTING (Passerina ciris) [b]
Icteridae (Troupials and Allies)
RED-WINGED BLACKBIRD (Agelaius phoeniceus)
EASTERN MEADOWLARK (Sturnella magna)
RED-BREASTED MEADOWLARK (Sturnella militaris)
MELODIOUS BLACKBIRD (Dives dives)
GREAT-TAILED GRACKLE (Quiscalus mexicanus)

A Yellow-throated Euphonia shows nicely his namesake body part. Photo by participant Paul Bisson.
SHINY COWBIRD (Molothrus bonariensis)
BRONZED COWBIRD (Molothrus aeneus)
GIANT COWBIRD (Molothrus oryzivorus)
BLACK-COWLED ORIOLE (Icterus prosthemelas)
BALTIMORE ORIOLE (Icterus galbula) [b]
SCARLET-RUMPED CACIQUE (SCARLET-RUMPED) (Cacicus uropygialis microrhynchus)
CHESTNUT-HEADED OROPENDOLA (Psarocolius wagleri) [N]
MONTEZUMA OROPENDOLA (Psarocolius montezuma) [N]
Fringillidae (Finches, Euphonias, and Allies)
SCRUB EUPHONIA (Euphonia affinis)
YELLOW-CROWNED EUPHONIA (Euphonia luteicapilla)
YELLOW-THROATED EUPHONIA (Euphonia hirundinacea)
ELEGANT EUPHONIA (Euphonia elegantissima)
SPOT-CROWNED EUPHONIA (Euphonia imitans)
OLIVE-BACKED EUPHONIA (Euphonia gouldi)
TAWNY-CAPPED EUPHONIA (Euphonia anneae)
GOLDEN-BROWED CHLOROPHONIA (Chlorophonia callophrys)
YELLOW-BELLIED SISKIN (Spinus xanthogastrus)
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus) [I]

For the full "tropical rainforest" experience, nothing beats a serenade from a Mantled Howler Monkey! Photo by participant Paul Bisson.
LONG-NOSED BAT (Rhynchonycteris naso)
NORTHERN GHOST BAT (Diclidurus albus)
COMMON TENT-MAKING BAT (Uroderma bilobatum)
MANTLED HOWLER MONKEY (Alouatta palliata)
WHITE-THROATED CAPUCHIN (Cebus capucinus)
CENTRAL AMERICAN SPIDER MONKEY (Ateles geoffroyi)
HOFFMANN'S TWO-TOED SLOTH (Choloepus hoffmanni)
BRAZILIAN RABBIT (Sylvilagus brasiliensis)
VARIEGATED SQUIRREL (Sciurus variegatoides)
RED-TAILED SQUIRREL (Sciurus granatensis)
ALFARO'S PYGMY SQUIRREL (Microsciurus alfari)
MEXICAN HAIRY PORCUPINE (Coendou mexicanus)
CENTRAL AMERICAN AGOUTI (Dasyprocta punctata)
GRAY FOX (Urocyon cinereoargenteus)
NORTHERN RACCOON (Procyon lotor)
WHITE-NOSED COATI (Nasua narica)
COLLARED PECCARY (Tayassu tajacu)

The Helmeted Basilisk (also known as Helmeted Iguana), is able to adjust its body color to help with thermoregulation. Photo by participant Paul Bisson.
Herps
SLENDER ANOLE (Anolis limifrons)
GREEN IGUANA (Iguana iguana)
BLACK SPINY-TAILED IGUANA (Ctenosaura similis)
COMMON BASILISK (Basiliscus basiliscus)
GREEN BASILISK (Basiliscus plumifrons)
STRIPED BASILISK (Basiliscus vittatus)
HELMETED BASILISK (Corytophanes cristatus)
TROPICAL HOUSE GECKO (Hemidactylus mabouia)
YELLOW-SPOTTED NIGHT LIZARD (Lepidophyma flavimaculatum)
CENTRAL AMERICAN WHIPTAIL (Ameiva festiva)
HIGHLAND ALLIGATOR LIZARD (Mesaspis monticola)
GREEN SPINY LIZARD (Sceloporus malachiticus)
GREEN PARROT SNAKE (Leptophis ahaetulla)
BLUNT-HEADED TREE SNAKE (Imantodes cenchoa)
AMERICAN CROCODILE (Crocodylus acutus)
SPECTACLED CAIMAN (Caiman crocodilus)
SMOKY JUNGLE FROG (Leptodactylus pentadactylus)
FORRER'S LEOPARD FROG (Lithobates forreri)
STRAWBERRY POISON DART FROG (Dendrobates pumilio)
SMOOTH-SKINNED TOAD (Bufo haematicus)
CANE TOAD (Rhinella marina)
Totals for the tour: 489 bird taxa and 17 mammal taxa