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This tour targets some of the unique, off-the-beaten-path birds that you might not find on a typical tour of central Costa Rica. The rare Yellow-billed Cotinga certainly qualifies, and was a major highlight of this year's tour. Photo by guide Tom Johnson.
From the comfortable breezes of the Central Valley to the misty heights of the Cerro de la Muerte and the steamy rainforests of the southern Pacific coast, this tour really packed in a LOT of habitat and elevational diversity into 8 full days in Costa Rica. Some participants stayed on for Part II in the northern reaches of the country.
After arriving from the airport in San Jose, we began with a breezy evening of birding at the Hotel Bougainvillea in Santo Domingo de Heredia, finding White-tailed Kite, Rufous-naped Wren, and Crimson-fronted Parakeets.
Our first full day of the tour was primarily a travel day. We birded in several spots in the montane forests of Cerro de la Muerte, but this was mostly in passing during our driving to Talari Mountain Lodge near San Isidro. While our endeavors were a bit hampered by light rain and fog, we found a great mixed flock along the entrance road to Paraiso Quetzal -- here we had Flame-throated Warblers, Yellow-winged Vireo, and Black-thighed Grosbeaks. We had a hard time focusing on our nice lunch at the Paraiso Quetzal lodge because of all of the hummingbirds zipping around outside. Fiery-throated Hummingbirds, common here, stole the show. We descended from the chilly highlands to our warmer lodging in the forest at Talari Mountain Lodge, where Fiery-billed Aracaris and a large flock of swifts helped to put the day to bed.
The next day found us exploring Alexander Skutch's estate at Los Cusingos. Charming Hummingbird, skulking Great Tinamous, Rufous Piha, and a plethora of tanagers met us in the hill forest here. We returned to Talari Mountain Lodge for the afternoon, where a watch along the river sent Peregrine Falcon and Lineated Woodpecker our way for great views. We capped the evening off with a Tropical Screech-Owl just outside our rooms.
After a morning walk on the trails at Talari turned up White-winged Becard, Red-crowned Ant-Tanager, and others, we hopped back on the bus and headed to the hills along the coast, to the steamy but very impressive Esquinas Rainforest Lodge. Birding the open rice fields along the entrance road was very exciting, with Purple Gallinules everywhere, Crested Guan and Black-mandibled Toucan perched up in trees, and Costa Rican Swifts and Muscovy Ducks flying by. In a short period of time before dinner, we found about 70 species in the open habitats along the entrance road.
The next day, we awoke to the whistled song of the very local Black-cheeked Ant-Tanager outside our rooms while Great Curassows paraded around on the grounds and a Baird's Trogon sang endlessly from the canopy. We headed to the Golfito Road, where the track through the forest gave us the opportunity to see several endemic Black-cheeked Ant-Tanagers very well at close range. Other highlights along the road included White-necked Puffbird and a soaring King Vulture. In Golfito, we enjoyed shorebirds on the mudflats and especially the Scarlet Macaws munching on almonds near the town center. A return to Esquinas Rainforest Lodge gave us time to again peruse the rice fields along the entrance road, this time turning up an adult Gray-lined Hawk among other goodies.
A very early morning drive the next day took us to the Rincon Bridge at the base of the Osa Peninsula. Here, we delighted in oodles of stunning cotingas, including at least 18 Yellow-billed Cotingas and 3 male Turquoise Cotingas -- wow! Additionally, Gray-headed Kites circled overhead, Boat-billed Herons perched along the river, and a Cinnamon Becard delighted in the forested edge.
After our final night at Esquinas Rainforest Lodge, we struck out across the hot lowlands toward the Panama border, stopping en route to bird at the Coto 47 marsh. Here we found such recent additions to the Costa Rican avifauna as Veraguan Mango and Rusty-margined Flycatcher and enjoyed low views of Lesser Yellow-headed Vulture as well. Our ascent toward San Vito led us to Wilson Botanical Gardens, our home for two nights.
Wilson offered plenty of options during our two days there. Some enjoyed the hummingbirds best -- we found a few remarkable Purple-crowned Fairies here; others took delight in the flocks of forest birds, including the charismatic Scale-crested Pygmy-Tyrant and the stealthy Black-faced Antthrush; night birding here turned up a most cooperative Mottled Owl as well.
The final day found us traveling back to San Jose. We took a route along the Pacific Coast, with a lovely stop at the Rio Tarcoles in the afternoon. Here we admired huge American Crocodiles in the river and also saw Turquoise-browed Motmot and Bat Falcons, among others. A bit of a struggle through evening traffic in San Jose took us back to the Hotel Bougainvillea, where we enjoyed a delightful dinner and toasted our good birding fortune for the week.
We had a great group, and whether this was a first time in the tropics or an opportunity to find a few out-of-the-way endemics, I think everyone enjoyed their time birding the south of Costa Rica. For Jay and myself, thank you all, and pura vida!
-Tom Johnson
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Tinamidae (Tinamous)
GREAT TINAMOU (Tinamus major)
LITTLE TINAMOU (Crypturellus soui) [*]
Anatidae (Ducks, Geese, and Waterfowl)
BLACK-BELLIED WHISTLING-DUCK (Dendrocygna autumnalis)
MUSCOVY DUCK (Cairina moschata)
BLUE-WINGED TEAL (Anas discors)
Cracidae (Guans, Chachalacas, and Curassows)
GRAY-HEADED CHACHALACA (Ortalis cinereiceps)
CRESTED GUAN (Penelope purpurascens)

Lesser Yellow-headed Vultures often fly quite low, allowing for close study of their colorful head skin. Photo by guide Tom Johnson.
BLACK GUAN (Chamaepetes unicolor) GREAT CURASSOW (Crax rubra)
Odontophoridae (New World Quail)
MARBLED WOOD-QUAIL (Odontophorus gujanensis) [*]
Ciconiidae (Storks)
WOOD STORK (Mycteria americana)
Fregatidae (Frigatebirds)
MAGNIFICENT FRIGATEBIRD (Fregata magnificens)
Phalacrocoracidae (Cormorants and Shags)
NEOTROPIC CORMORANT (Phalacrocorax brasilianus)
Anhingidae (Anhingas)
ANHINGA (Anhinga anhinga)
Pelecanidae (Pelicans)
BROWN PELICAN (Pelecanus occidentalis)
Ardeidae (Herons, Egrets, and Bitterns)
BARE-THROATED TIGER-HERON (Tigrisoma mexicanum)
GREAT EGRET (Ardea alba)

This Tropical Screech-Owl was a nice bonus feature of our excellent accommodations at Hotel Bougainvillea. Photo by guide Tom Johnson.
SNOWY EGRET (Egretta thula) LITTLE BLUE HERON (Egretta caerulea)
TRICOLORED HERON (Egretta tricolor)
CATTLE EGRET (Bubulcus ibis)
GREEN HERON (Butorides virescens)
BOAT-BILLED HERON (Cochlearius cochlearius)
Threskiornithidae (Ibises and Spoonbills)
WHITE IBIS (Eudocimus albus)
ROSEATE SPOONBILL (Platalea ajaja)
Cathartidae (New World Vultures)
BLACK VULTURE (Coragyps atratus)
TURKEY VULTURE (Cathartes aura)
LESSER YELLOW-HEADED VULTURE (Cathartes burrovianus)
KING VULTURE (Sarcoramphus papa)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus)
Accipitridae (Hawks, Eagles, and Kites)
PEARL KITE (Gampsonyx swainsonii)
WHITE-TAILED KITE (Elanus leucurus)
GRAY-HEADED KITE (Leptodon cayanensis)
SWALLOW-TAILED KITE (Elanoides forficatus)
BLACK HAWK-EAGLE (Spizaetus tyrannus)
DOUBLE-TOOTHED KITE (Harpagus bidentatus)

This Band-tailed Barbthroat was perched low along the Golfito Rd., providing our only study of the species for the tour. Photo by guide Tom Johnson.
COMMON BLACK HAWK (MANGROVE) (Buteogallus anthracinus subtilis) ROADSIDE HAWK (Rupornis magnirostris)
BROAD-WINGED HAWK (Buteo platypterus)
GRAY HAWK (Buteo plagiatus)
GRAY-LINED HAWK (Buteo nitidus)
SHORT-TAILED HAWK (Buteo brachyurus)
Rallidae (Rails, Gallinules, and Coots)
WHITE-THROATED CRAKE (Laterallus albigularis) [*]
GRAY-BREASTED CRAKE (Laterallus exilis) [*]
GRAY-NECKED WOOD-RAIL (Aramides cajaneus)
PURPLE GALLINULE (Porphyrio martinicus)
COMMON GALLINULE (Gallinula galeata)
Recurvirostridae (Stilts and Avocets)
BLACK-NECKED STILT (Himantopus mexicanus)
Charadriidae (Plovers and Lapwings)
BLACK-BELLIED PLOVER (Pluvialis squatarola)
SOUTHERN LAPWING (Vanellus chilensis)

Short bills sometimes equal shortcuts to nectar in the hummingbird world. This Purple-crowned Fairy attacked the base of red flowers as we watched near the canopy tower at Wilson Botanical Gardens. Photo by guide Tom Johnson.
WILSON'S PLOVER (Charadrius wilsonia) SEMIPALMATED PLOVER (Charadrius semipalmatus)
Jacanidae (Jacanas)
NORTHERN JACANA (Jacana spinosa)
Scolopacidae (Sandpipers and Allies)
SPOTTED SANDPIPER (Actitis macularius)
GREATER YELLOWLEGS (Tringa melanoleuca)
WILLET (Tringa semipalmata)
LESSER YELLOWLEGS (Tringa flavipes)
WHIMBREL (Numenius phaeopus)
RUDDY TURNSTONE (Arenaria interpres)
LEAST SANDPIPER (Calidris minutilla)
WESTERN SANDPIPER (Calidris mauri)
Laridae (Gulls, Terns, and Skimmers)
LAUGHING GULL (Leucophaeus atricilla)
ROYAL TERN (Thalasseus maximus)
SANDWICH TERN (Thalasseus sandvicensis)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
PALE-VENTED PIGEON (Patagioenas cayennensis)
SCALED PIGEON (Patagioenas speciosa)
RED-BILLED PIGEON (Patagioenas flavirostris)
RUDDY PIGEON (Patagioenas subvinacea)

A specialty of the highlands, Fiery-throated Hummingbird was perhaps the most colorful hummer of the tour. Photo by guide Tom Johnson.
SHORT-BILLED PIGEON (Patagioenas nigrirostris) INCA DOVE (Columbina inca)
RUDDY GROUND-DOVE (Columbina talpacoti)
BLUE GROUND-DOVE (Claravis pretiosa)
WHITE-TIPPED DOVE (Leptotila verreauxi)
GRAY-CHESTED DOVE (Leptotila cassinii)
WHITE-WINGED DOVE (Zenaida asiatica)
Cuculidae (Cuckoos)
SQUIRREL CUCKOO (Piaya cayana)
STRIPED CUCKOO (Tapera naevia)
SMOOTH-BILLED ANI (Crotophaga ani)
GROOVE-BILLED ANI (Crotophaga sulcirostris)
Strigidae (Owls)
TROPICAL SCREECH-OWL (Megascops choliba)
CRESTED OWL (Lophostrix cristata) [*]
SPECTACLED OWL (Pulsatrix perspicillata)
MOTTLED OWL (Ciccaba virgata)
Caprimulgidae (Nightjars and Allies)
LESSER NIGHTHAWK (Chordeiles acutipennis)

The namesake bird of Alexander Skutch's Los Cusingos estate, this Fiery-billed Aracari was perched outside the dining porch at Talari Mountain Lodge. Photo by guide Tom Johnson.
SHORT-TAILED NIGHTHAWK (Lurocalis semitorquatus) COMMON PAURAQUE (Nyctidromus albicollis)
Nyctibiidae (Potoos)
COMMON POTOO (Nyctibius griseus)
Apodidae (Swifts)
WHITE-CHINNED SWIFT (Cypseloides cryptus)
SPOT-FRONTED SWIFT (Cypseloides cherriei)
CHESTNUT-COLLARED SWIFT (Streptoprocne rutila)
WHITE-COLLARED SWIFT (Streptoprocne zonaris)
VAUX'S SWIFT (Chaetura vauxi)
COSTA RICAN SWIFT (Chaetura fumosa)
LESSER SWALLOW-TAILED SWIFT (Panyptila cayennensis)
Trochilidae (Hummingbirds)
WHITE-NECKED JACOBIN (Florisuga mellivora)
BAND-TAILED BARBTHROAT (Threnetes ruckeri)
GREEN HERMIT (Phaethornis guy)
LONG-BILLED HERMIT (Phaethornis longirostris)
STRIPE-THROATED HERMIT (Phaethornis striigularis)
GREEN VIOLETEAR (Colibri thalassinus)
PURPLE-CROWNED FAIRY (Heliothryx barroti)
VERAGUAN MANGO (Anthracothorax veraguensis)

These Black-mandibled Toucans were scouting for food along the entrance road near Esquinas Rainforest Lodge, much to the dismay of an excited group of songbirds. Photo by guide Tom Johnson.
MAGNIFICENT HUMMINGBIRD (Eugenes fulgens) LONG-BILLED STARTHROAT (Heliomaster longirostris)
FIERY-THROATED HUMMINGBIRD (Panterpe insignis)
VOLCANO HUMMINGBIRD (Selasphorus flammula)
GARDEN EMERALD (Chlorostilbon assimilis)
SCALY-BREASTED HUMMINGBIRD (Phaeochroa cuvierii)
CROWNED WOODNYMPH (Thalurania colombica)
WHITE-TAILED EMERALD (Elvira chionura)
CHARMING HUMMINGBIRD (Amazilia decora)
SNOWY-BELLIED HUMMINGBIRD (Amazilia edward)
RUFOUS-TAILED HUMMINGBIRD (Amazilia tzacatl)
BLUE-THROATED GOLDENTAIL (Hylocharis eliciae)
Trogonidae (Trogons)
SLATY-TAILED TROGON (Trogon massena)
BAIRD'S TROGON (Trogon bairdii)
GARTERED TROGON (Trogon caligatus)
BLACK-THROATED TROGON (Trogon rufus)
COLLARED TROGON (Trogon collaris)
Momotidae (Motmots)
BLUE-CROWNED MOTMOT (LESSON'S) (Momotus coeruliceps lessonii)
TURQUOISE-BROWED MOTMOT (Eumomota superciliosa)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)

Brown-throated Parakeets arrived in Costa Rica from Panama recently. These two were near Esquinas Rainforest Lodge. Photo by guide Tom Johnson.
AMAZON KINGFISHER (Chloroceryle amazona) GREEN KINGFISHER (Chloroceryle americana)
AMERICAN PYGMY KINGFISHER (Chloroceryle aenea)
Bucconidae (Puffbirds)
WHITE-NECKED PUFFBIRD (Notharchus hyperrhynchus)
Galbulidae (Jacamars)
RUFOUS-TAILED JACAMAR (Galbula ruficauda)
Ramphastidae (Toucans)
FIERY-BILLED ARACARI (Pteroglossus frantzii)
BLACK-MANDIBLED TOUCAN (CHESTNUT-MANDIBLED) (Ramphastos ambiguus swainsonii)
Picidae (Woodpeckers)
OLIVACEOUS PICULET (Picumnus olivaceus)
GOLDEN-NAPED WOODPECKER (Melanerpes chrysauchen)
RED-CROWNED WOODPECKER (Melanerpes rubricapillus)
HOFFMANN'S WOODPECKER (Melanerpes hoffmannii)
SMOKY-BROWN WOODPECKER (Picoides fumigatus)
RUFOUS-WINGED WOODPECKER (Piculus simplex)
GOLDEN-OLIVE WOODPECKER (Colaptes rubiginosus)
LINEATED WOODPECKER (Dryocopus lineatus)
PALE-BILLED WOODPECKER (Campephilus guatemalensis)
Falconidae (Falcons and Caracaras)
CRESTED CARACARA (Caracara cheriway)
YELLOW-HEADED CARACARA (Milvago chimachima)

This Scarlet Macaw was part of a small group eating almonds in downtown Golfito. Photo by guide Tom Johnson.
LAUGHING FALCON (Herpetotheres cachinnans) [*] BAT FALCON (Falco rufigularis)
PEREGRINE FALCON (Falco peregrinus)
Psittacidae (New World and African Parrots)
ORANGE-CHINNED PARAKEET (Brotogeris jugularis)
BROWN-HOODED PARROT (Pyrilia haematotis)
BLUE-HEADED PARROT (Pionus menstruus)
WHITE-CROWNED PARROT (Pionus senilis)
RED-LORED PARROT (Amazona autumnalis)
YELLOW-NAPED PARROT (Amazona auropalliata)
WHITE-FRONTED PARROT (Amazona albifrons)
MEALY PARROT (Amazona farinosa)
BROWN-THROATED PARAKEET (Eupsittula pertinax)
SCARLET MACAW (Ara macao)
CRIMSON-FRONTED PARAKEET (Psittacara finschi)
Thamnophilidae (Typical Antbirds)
BLACK-HOODED ANTSHRIKE (Thamnophilus bridgesi)
RUSSET ANTSHRIKE (Thamnistes anabatinus)
PLAIN ANTVIREO (Dysithamnus mentalis)
SLATY ANTWREN (Myrmotherula schisticolor)

This gorgeous Black-striped Woodcreeper sat up near the lodge at Esquinas Rainforest Lodge as we were about to leave. Photo by guide Tom Johnson.
DOT-WINGED ANTWREN (Microrhopias quixensis) DUSKY ANTBIRD (Cercomacra tyrannina)
CHESTNUT-BACKED ANTBIRD (Myrmeciza exsul)
Grallariidae (Antpittas)
STREAK-CHESTED ANTPITTA (Hylopezus perspicillatus) [*]
Formicariidae (Antthrushes)
BLACK-FACED ANTTHRUSH (Formicarius analis)
Furnariidae (Ovenbirds and Woodcreepers)
OLIVACEOUS WOODCREEPER (Sittasomus griseicapillus)
WEDGE-BILLED WOODCREEPER (Glyphorynchus spirurus)
NORTHERN BARRED-WOODCREEPER (Dendrocolaptes sanctithomae)
COCOA WOODCREEPER (Xiphorhynchus susurrans)
BLACK-STRIPED WOODCREEPER (Xiphorhynchus lachrymosus)
SPOTTED WOODCREEPER (Xiphorhynchus erythropygius)
STREAK-HEADED WOODCREEPER (Lepidocolaptes souleyetii)
PLAIN XENOPS (Xenops minutus)
BUFF-THROATED FOLIAGE-GLEANER (EXSERTUS) (Automolus ochrolaemus exsertus)
RUDDY TREERUNNER (Margarornis rubiginosus)
RED-FACED SPINETAIL (Cranioleuca erythrops)
PALE-BREASTED SPINETAIL (Synallaxis albescens)
Tyrannidae (Tyrant Flycatchers)
SOUTHERN BEARDLESS-TYRANNULET (Camptostoma obsoletum)
YELLOW TYRANNULET (Capsiempis flaveola)
GREENISH ELAENIA (Myiopagis viridicata)
YELLOW-BELLIED ELAENIA (Elaenia flavogaster)
OLIVE-STRIPED FLYCATCHER (Mionectes olivaceus)
OCHRE-BELLIED FLYCATCHER (Mionectes oleagineus)

This Scale-crested Pygmy-Tyrant showed off its fancy crown feathers along the forest trails at Wilson Botanical Gardens. Photo by guide Tom Johnson.
PALTRY TYRANNULET (Zimmerius vilissimus) SCALE-CRESTED PYGMY-TYRANT (Lophotriccus pileatus)
NORTHERN BENTBILL (Oncostoma cinereigulare)
SLATE-HEADED TODY-FLYCATCHER (Poecilotriccus sylvia)
COMMON TODY-FLYCATCHER (Todirostrum cinereum)
EYE-RINGED FLATBILL (Rhynchocyclus brevirostris)
YELLOW-OLIVE FLYCATCHER (Tolmomyias sulphurescens)
RUDDY-TAILED FLYCATCHER (Terenotriccus erythrurus)
SULPHUR-RUMPED FLYCATCHER (Myiobius sulphureipygius aureatus)
BRAN-COLORED FLYCATCHER (Myiophobus fasciatus)
OLIVE-SIDED FLYCATCHER (Contopus cooperi)
WESTERN WOOD-PEWEE (Contopus sordidulus)
YELLOW-BELLIED FLYCATCHER (Empidonax flaviventris)
ACADIAN FLYCATCHER (Empidonax virescens)
WILLOW FLYCATCHER (Empidonax traillii)
BLACK-CAPPED FLYCATCHER (Empidonax atriceps)
BLACK PHOEBE (Sayornis nigricans)
BRIGHT-RUMPED ATTILA (Attila spadiceus)
RUFOUS MOURNER (Rhytipterna holerythra)
DUSKY-CAPPED FLYCATCHER (Myiarchus tuberculifer)
GREAT KISKADEE (Pitangus sulphuratus)
BOAT-BILLED FLYCATCHER (Megarynchus pitangua)
RUSTY-MARGINED FLYCATCHER (Myiozetetes cayanensis)
SOCIAL FLYCATCHER (Myiozetetes similis)
GRAY-CAPPED FLYCATCHER (Myiozetetes granadensis)
STREAKED FLYCATCHER (Myiodynastes maculatus)
PIRATIC FLYCATCHER (Legatus leucophaius)
TROPICAL KINGBIRD (Tyrannus melancholicus)

So nice, we featured it twice; the 18 Yellow-billed Cotingas that we saw at Rincon Bridge on the Osa Peninsula were some of the rarest birds we saw during the tour. Photo by guide Tom Johnson.
SCISSOR-TAILED FLYCATCHER (Tyrannus forficatus) FORK-TAILED FLYCATCHER (Tyrannus savana)
Cotingidae (Cotingas)
TURQUOISE COTINGA (Cotinga ridgwayi)
RUFOUS PIHA (Lipaugus unirufus)
YELLOW-BILLED COTINGA (Carpodectes antoniae)
Pipridae (Manakins)
WHITE-RUFFED MANAKIN (Corapipo altera)
BLUE-CROWNED MANAKIN (Lepidothrix coronata)
ORANGE-COLLARED MANAKIN (Manacus aurantiacus)
RED-CAPPED MANAKIN (Ceratopipra mentalis)
Tityridae (Tityras and Allies)
BLACK-CROWNED TITYRA (Tityra inquisitor)
MASKED TITYRA (Tityra semifasciata)
CINNAMON BECARD (Pachyramphus cinnamomeus)
WHITE-WINGED BECARD (Pachyramphus polychopterus)
ROSE-THROATED BECARD (Pachyramphus aglaiae)
Vireonidae (Vireos, Shrike-Babblers, and Erpornis)
YELLOW-THROATED VIREO (Vireo flavifrons)
YELLOW-WINGED VIREO (Vireo carmioli)
LESSER GREENLET (Hylophilus decurtatus)
GREEN SHRIKE-VIREO (Vireolanius pulchellus) [*]
RUFOUS-BROWED PEPPERSHRIKE (Cyclarhis gujanensis)
Corvidae (Crows, Jays, and Magpies)
BROWN JAY (Psilorhinus morio)
Hirundinidae (Swallows)
BLUE-AND-WHITE SWALLOW (Pygochelidon cyanoleuca)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)

This Flame-throated Warbler, a highland specialty of western Panama and Costa Rica, was in a busy mixed flock on the Cerro de la Muerte. Photo by guide Tom Johnson.
GRAY-BREASTED MARTIN (Progne chalybea) MANGROVE SWALLOW (Tachycineta albilinea)
BARN SWALLOW (Hirundo rustica)
Troglodytidae (Wrens)
HOUSE WREN (Troglodytes aedon)
OCHRACEOUS WREN (Troglodytes ochraceus)
TIMBERLINE WREN (Thryorchilus browni)
RUFOUS-NAPED WREN (Campylorhynchus rufinucha)
BLACK-BELLIED WREN (Pheugopedius fasciatoventris)
RUFOUS-BREASTED WREN (Pheugopedius rutilus)
PLAIN WREN (Cantorchilus modestus)
RIVERSIDE WREN (Cantorchilus semibadius)
WHITE-BREASTED WOOD-WREN (Henicorhina leucosticta) [*]
GRAY-BREASTED WOOD-WREN (Henicorhina leucophrys)
Polioptilidae (Gnatcatchers)
LONG-BILLED GNATWREN (Ramphocaenus melanurus)
TROPICAL GNATCATCHER (Polioptila plumbea)
Turdidae (Thrushes and Allies)
BLACK-BILLED NIGHTINGALE-THRUSH (Catharus gracilirostris)
ORANGE-BILLED NIGHTINGALE-THRUSH (Catharus aurantiirostris)
SOOTY THRUSH (Turdus nigrescens)
CLAY-COLORED THRUSH (Turdus grayi)
WHITE-THROATED THRUSH (Turdus assimilis)
Mimidae (Mockingbirds and Thrashers)
TROPICAL MOCKINGBIRD (Mimus gilvus)
Parulidae (New World Warblers)
NORTHERN WATERTHRUSH (Parkesia noveboracensis)

This Speckled Tanager was interested in the delicious bananas on the veranda at Wilson Botanical Gardens. Photo by guide Tom Johnson.
GOLDEN-WINGED WARBLER (Vermivora chrysoptera) BLACK-AND-WHITE WARBLER (Mniotilta varia)
FLAME-THROATED WARBLER (Oreothlypis gutturalis)
TENNESSEE WARBLER (Oreothlypis peregrina)
MASKED YELLOWTHROAT (CHIRIQU) (Geothlypis aequinoctialis chiriquensis)
AMERICAN REDSTART (Setophaga ruticilla)
TROPICAL PARULA (Setophaga pitiayumi)
YELLOW WARBLER (Setophaga petechia)
CHESTNUT-SIDED WARBLER (Setophaga pensylvanica)
BLACK-THROATED GREEN WARBLER (Setophaga virens)
RUFOUS-CAPPED WARBLER (Basileuterus rufifrons)
BLACK-CHEEKED WARBLER (Basileuterus melanogenys)
BUFF-RUMPED WARBLER (Myiothlypis fulvicauda)
WILSON'S WARBLER (Cardellina pusilla)
SLATE-THROATED REDSTART (Myioborus miniatus)
COLLARED REDSTART (Myioborus torquatus)
WRENTHRUSH (Zeledonia coronata) [*]
Thraupidae (Tanagers and Allies)
GRAY-HEADED TANAGER (Eucometis penicillata)
CHERRIE'S TANAGER (Ramphocelus costaricensis)
BLUE-GRAY TANAGER (Thraupis episcopus)
PALM TANAGER (Thraupis palmarum)
GOLDEN-HOODED TANAGER (Tangara larvata)

Volcano Juncos are endemic to the highlands of Costa Rica and western Panama. Photo by guide Tom Johnson.
SPECKLED TANAGER (Tangara guttata) SPANGLE-CHEEKED TANAGER (Tangara dowii)
BAY-HEADED TANAGER (Tangara gyrola)
SILVER-THROATED TANAGER (Tangara icterocephala)
SCARLET-THIGHED DACNIS (Dacnis venusta)
BLUE DACNIS (Dacnis cayana)
SHINING HONEYCREEPER (Cyanerpes lucidus)
RED-LEGGED HONEYCREEPER (Cyanerpes cyaneus)
GREEN HONEYCREEPER (Chlorophanes spiza)
SLATY FLOWERPIERCER (Diglossa plumbea)
BLUE-BLACK GRASSQUIT (Volatinia jacarina)
RUDDY-BREASTED SEEDEATER (Sporophila minuta)
THICK-BILLED SEED-FINCH (Sporophila funerea)
VARIABLE SEEDEATER (Sporophila corvina)
WHITE-COLLARED SEEDEATER (Sporophila torqueola)
YELLOW-BELLIED SEEDEATER (Sporophila nigricollis)
SLATE-COLORED SEEDEATER (Sporophila schistacea)
BANANAQUIT (Coereba flaveola)
YELLOW-FACED GRASSQUIT (Tiaris olivaceus)
BUFF-THROATED SALTATOR (Saltator maximus)

This Black-cheeked Ant-Tanager was one of a small group of this localized Costa Rican endemic that we saw along the Golfito Road. Photo by guide Tom Johnson.
GRAYISH SALTATOR (Saltator coerulescens) STREAKED SALTATOR (Saltator striatipectus)
Emberizidae (Buntings and New World Sparrows)
YELLOW-THIGHED FINCH (Pselliophorus tibialis)
LARGE-FOOTED FINCH (Pezopetes capitalis)
ORANGE-BILLED SPARROW (Arremon aurantiirostris)
COSTA RICAN BRUSH-FINCH (Arremon costaricensis)
BLACK-STRIPED SPARROW (Arremonops conirostris)
RUFOUS-COLLARED SPARROW (Zonotrichia capensis)
VOLCANO JUNCO (Junco vulcani)
COMMON CHLOROSPINGUS (Chlorospingus flavopectus)
SOOTY-CAPPED CHLOROSPINGUS (Chlorospingus pileatus)
Cardinalidae (Cardinals and Allies)
SUMMER TANAGER (Piranga rubra)
WHITE-WINGED TANAGER (Piranga leucoptera)
RED-CROWNED ANT-TANAGER (Habia rubica)
BLACK-CHEEKED ANT-TANAGER (Habia atrimaxillaris) [E]
BLACK-THIGHED GROSBEAK (Pheucticus tibialis)
ROSE-BREASTED GROSBEAK (Pheucticus ludovicianus)
Icteridae (Troupials and Allies)
EASTERN MEADOWLARK (Sturnella magna)
RED-BREASTED BLACKBIRD (Sturnella militaris)
MELODIOUS BLACKBIRD (Dives dives)
GREAT-TAILED GRACKLE (Quiscalus mexicanus)
SHINY COWBIRD (Molothrus bonariensis)
BRONZED COWBIRD (Molothrus aeneus)
GIANT COWBIRD (Molothrus oryzivorus)
ORCHARD ORIOLE (Icterus spurius)
BALTIMORE ORIOLE (Icterus galbula)

Male Euphonias are snazzy, but even the females can be well-marked. This female Spot-crowned Euphonia was at Wilson Botanical Gardens. Photo by guide Tom Johnson.
SCARLET-RUMPED CACIQUE (SCARLET-RUMPED) (Cacicus uropygialis microrhynchus) CRESTED OROPENDOLA (Psarocolius decumanus)
Fringillidae (Finches, Euphonias, and Allies)
YELLOW-CROWNED EUPHONIA (Euphonia luteicapilla)
THICK-BILLED EUPHONIA (Euphonia laniirostris)
SPOT-CROWNED EUPHONIA (Euphonia imitans)
LESSER GOLDFINCH (Spinus psaltria)
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus) [I]
LONG-NOSED BAT (Rhynchonycteris naso)
GREATER WHITE-LINED BAT (Saccopteryx bilineata)
JAMAICAN FRUIT-EATING BAT (Artibeus jamaicensis)
WHITE-THROATED CAPUCHIN (Cebus capucinus)
VARIEGATED SQUIRREL (Sciurus variegatoides)
RED-TAILED SQUIRREL (Sciurus granatensis)
MEXICAN HAIRY PORCUPINE (Coendou mexicanus)
CENTRAL AMERICAN AGOUTI (Dasyprocta punctata)
KINKAJOU (Potos flavus)
Herps
GREEN IGUANA (Iguana iguana)

American Crocodiles smiled at us from the sandbars below the Rio Tarcoles bridge on the last day of the tour. Photo by guide Tom Johnson.
COMMON BASILISK (Basiliscus basiliscus) TROPICAL HOUSE GECKO (Hemidactylus mabouia)
BOA CONSTRICTOR (Boa constrictor)
AMERICAN CROCODILE (Crocodylus acutus)
SPECTACLED CAIMAN (Caiman crocodilus)
CANE TOAD (Bufo marinus)
BRILLIANT FOREST FROG (Rana warszewitschii)
MESO-AMERICAN SLIDER (Trachemys venusta)
Totals for the tour: 329 bird taxa and 9 mammal taxa