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The view from Arenal Observatory Lodge is simply breathtaking. Here, Lake Arenal is in the background on the left and the slope of the Arenal Volcano is on the right -- and in the middle, forest with LOTS of birds. Photo by guide Cory Gregory.
This wonderful loop through Northern Costa Rica took us from the cool Central Valley into the misty, breezy middle elevations and highlands of some of the country's famous volcanoes, with a nice warm interlude in the Lake Nicaragua lowlands around Caño Negro. Weather was very pleasant throughout, with very little rain during our birding outings.
We started off with a morning on Volcan Poás, finding highland specialities like Golden-browed Chlorophonias feeding low on a fruiting fig tree. At lunchtime, we enjoyed the hummingbirds of the La Paz Waterfall Gardens, and as a nice encore, we stopped at some roadside feeders that had very close Red-headed Barbets, Emerald Toucanets, tanagers, and orioles gorging themselves on bananas.
Our home for two nights in mid-elevation forest was Hotel Bosque de Paz, a lovely spot indeed. We enjoyed the Black Guans, Chestnut-capped Brushfinches, and strange Pacas at the feeders as well as the Brown-billed Scythebills, Long-tailed Silky-flycatchers, and nightingale-thrushes in the forest.
Dropping in elevation and increasing in temperature, we shifted north to the lowlands along the Nicaraguan border. We added a boat trip in the Medio Queso wetland this year, a great way for us to see some wonderful birds like Pinnated Bittern (two, and CLOSE!), American Pygmy-Kingfisher, Nicaraguan Grackle (we missed it at Caño Negro, so we were pleased to find it here), and scores of Black-bellied Whistling-Ducks. On the road to Caño Negro, we found Nicaraguan Seed-Finch, Double-striped Thick-knee, and at night, Great Potoo (WOW! They were close and marvelous!), Striped Owl, and Black-and-white Owl. Our boat trips into the flooded Caño Negro were very interesting, allowing us some close views at birds like Sungrebe, Spot-breasted Wren, Gray-headed Kite, Black-collared Hawk, Slaty-tailed Trogon, and Olive-crowned Yellowthroat. Water levels were still high after flooding in late November 2016 due to the devastating Hurricane Otto.
Our final stay on the tour was at the always wonderful (and always surprising!) Arenal Observatory Lodge at the foot of the Arenal Volcano. A fruiting fig tree in the gardens was erupting, taking the place of the now-quiet volcano. We saw tons of tanagers, toucans, thrushes, and Crested Guans eating the fruit here -- we even spotted a rare Yellow-eared Toucanet early on our second morning. The forests of Arenal were very productive, and we found some fun birds including White-throated Shrike-Tanager, Orange-bellied Trogon, Song Wren, Ocellated Antbird, Olive-backed Euphonia, Slate-colored Grosbeak, Thicket Antpitta, and more. By Lake Arenal, we were able to compare the scarce Keel-billed and Broad-billed motmots in an area where they sometimes pair up together, and Cory spotted a surprising Northern Tamandua hustling down a tree and then slowly climbing a hillside across from us.
Cory and I were delighted to share this piece of Costa Rica with you - we really saw a lot of this fantastic country in a week, with plenty of highlights (birds and otherwise). Thank you for birding with us, and we hope to see you out on the trails again soon!
Special thanks go to Vernon Campos for driving and helping us as an outstanding third guide on this fun and exciting tour.
Cheers,
Tom
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Anatidae (Ducks, Geese, and Waterfowl)
BLACK-BELLIED WHISTLING-DUCK (Dendrocygna autumnalis)
BLUE-WINGED TEAL (Anas discors)
Cracidae (Guans, Chachalacas, and Curassows)
CRESTED GUAN (Penelope purpurascens)

We had some splendid views of this Brown-billed Scythebill in the forest near Bosque de Paz. This species has to lean back just to fit itself on a tree trunk! Photo by guide Tom Johnson.
BLACK GUAN (Chamaepetes unicolor)
GREAT CURASSOW (Crax rubra)
Ciconiidae (Storks)
JABIRU (Jabiru mycteria)
WOOD STORK (Mycteria americana)
Phalacrocoracidae (Cormorants and Shags)
NEOTROPIC CORMORANT (Phalacrocorax brasilianus)
Anhingidae (Anhingas)
ANHINGA (Anhinga anhinga)
Ardeidae (Herons, Egrets, and Bitterns)
PINNATED BITTERN (Botaurus pinnatus)
FASCIATED TIGER-HERON (Tigrisoma fasciatum)
BARE-THROATED TIGER-HERON (Tigrisoma mexicanum)
GREAT BLUE HERON (Ardea herodias)

Fruit feeders provide some great opportunities to study frugivores up close. This Emerald Toucanet was one of several coming in to bananas at Cinchona. Photo by guide Cory Gregory.
GREAT EGRET (Ardea alba)
SNOWY EGRET (Egretta thula)
LITTLE BLUE HERON (Egretta caerulea)
TRICOLORED HERON (Egretta tricolor)
CATTLE EGRET (Bubulcus ibis)
GREEN HERON (Butorides virescens)
BLACK-CROWNED NIGHT-HERON (Nycticorax nycticorax)
Threskiornithidae (Ibises and Spoonbills)
WHITE IBIS (Eudocimus albus)
GLOSSY IBIS (Plegadis falcinellus)

A new species as of 2016 (after the split of Gray-necked Wood-Rail), this Russet-naped Wood-Rail was a great addition to our trip near Caño Negro. Photo by guide Tom Johnson.
GREEN IBIS (Mesembrinibis cayennensis)
ROSEATE SPOONBILL (Platalea ajaja)
Cathartidae (New World Vultures)
BLACK VULTURE (Coragyps atratus)
TURKEY VULTURE (Cathartes aura)
LESSER YELLOW-HEADED VULTURE (Cathartes burrovianus)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus)
Accipitridae (Hawks, Eagles, and Kites)
WHITE-TAILED KITE (Elanus leucurus)
GRAY-HEADED KITE (Leptodon cayanensis)
ORNATE HAWK-EAGLE (Spizaetus ornatus) [*]
BLACK-COLLARED HAWK (Busarellus nigricollis)

A Fasciated Tiger-Heron showed off repeatedly along the entrance drive to Arenal. Photo by guide Tom Johnson.
SNAIL KITE (Rostrhamus sociabilis)
NORTHERN HARRIER (Circus cyaneus)
SHARP-SHINNED HAWK (Accipiter striatus)
ROADSIDE HAWK (Rupornis magnirostris)
HARRIS'S HAWK (Parabuteo unicinctus)
WHITE HAWK (Pseudastur albicollis)
GRAY HAWK (Buteo plagiatus)
BROAD-WINGED HAWK (Buteo platypterus)
SHORT-TAILED HAWK (Buteo brachyurus)
RED-TAILED HAWK (Buteo jamaicensis)
Rallidae (Rails, Gallinules, and Coots)
WHITE-THROATED CRAKE (Laterallus albigularis)

Army ants had this Yellowish Flycatcher distracted at Bosque de Paz. Photo by guide Cory Gregory.
RUSSET-NAPED WOOD-RAIL (Aramides albiventris)
PURPLE GALLINULE (Porphyrio martinicus)
Heliornithidae (Finfoots)
SUNGREBE (Heliornis fulica)
Burhinidae (Thick-knees)
DOUBLE-STRIPED THICK-KNEE (Burhinus bistriatus)
Recurvirostridae (Stilts and Avocets)
BLACK-NECKED STILT (Himantopus mexicanus)
Jacanidae (Jacanas)
NORTHERN JACANA (Jacana spinosa)
Scolopacidae (Sandpipers and Allies)
LEAST SANDPIPER (Calidris minutilla)

As we were leaving Caño Negro, this Streak-headed Woodcreeper put on a good show next to the bus. Photo by guide Tom Johnson.
WILSON'S SNIPE (Gallinago delicata)
SPOTTED SANDPIPER (Actitis macularius)
SOLITARY SANDPIPER (Tringa solitaria)
GREATER YELLOWLEGS (Tringa melanoleuca)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
PALE-VENTED PIGEON (Patagioenas cayennensis)
SCALED PIGEON (Patagioenas speciosa)
RED-BILLED PIGEON (Patagioenas flavirostris)

We were pretty stoked to find this Yellow-eared Toucanet in the fruiting fig trees at Arenal Observatory Lodge. Photo by guide Tom Johnson.
BAND-TAILED PIGEON (Patagioenas fasciata)
INCA DOVE (Columbina inca)
PLAIN-BREASTED GROUND-DOVE (Columbina minuta)
RUDDY GROUND-DOVE (Columbina talpacoti)
BLUE GROUND-DOVE (Claravis pretiosa)
WHITE-TIPPED DOVE (Leptotila verreauxi)
GRAY-CHESTED DOVE (Leptotila cassinii)
GRAY-HEADED DOVE (Leptotila plumbeiceps)
QUAIL-DOVE SP. (Columbidae sp. )

Our boat trips on the Caño Negro were great for all kinds of wildlife, not just birds. This Spectacled Caiman gave us a nice grin as we cruised past. Photo by guide Cory Gregory.
WHITE-WINGED DOVE (Zenaida asiatica)
Cuculidae (Cuckoos)
GROOVE-BILLED ANI (Crotophaga sulcirostris)
SQUIRREL CUCKOO (Piaya cayana)
MANGROVE CUCKOO (Coccyzus minor)
Tytonidae (Barn-Owls)
BARN OWL (Tyto alba)
Strigidae (Owls)
PACIFIC SCREECH-OWL (Megascops cooperi)
TROPICAL SCREECH-OWL (Megascops choliba)
FERRUGINOUS PYGMY-OWL (Glaucidium brasilianum)
BLACK-AND-WHITE OWL (Ciccaba nigrolineata)
STRIPED OWL (Pseudoscops clamator)
Caprimulgidae (Nightjars and Allies)
LESSER NIGHTHAWK (Chordeiles acutipennis)

We saw this male Tawny-capped Euphonia really well near the river at Arenal Observatory Lodge. Photo by guide Tom Johnson.
COMMON PAURAQUE (Nyctidromus albicollis)
Nyctibiidae (Potoos)
GREAT POTOO (Nyctibius grandis)
Apodidae (Swifts)
WHITE-COLLARED SWIFT (Streptoprocne zonaris)
GRAY-RUMPED SWIFT (Chaetura cinereiventris)
Trochilidae (Hummingbirds)
WHITE-NECKED JACOBIN (Florisuga mellivora)
GREEN HERMIT (Phaethornis guy)
STRIPE-THROATED HERMIT (Phaethornis striigularis)

Several species of hummingbirds patronized the flowers in the gardens at Arenal Observatory Lodge. This Violet-headed Hummingbird shows off its blue-purple forehead patch and short bill. Photo by guide Cory Gregory.
PURPLE-CROWNED FAIRY (Heliothryx barroti)
GREEN-BREASTED MANGO (Anthracothorax prevostii)
GREEN THORNTAIL (Discosura conversii)
BLACK-CRESTED COQUETTE (Lophornis helenae)
GREEN-CROWNED BRILLIANT (Heliodoxa jacula)
MAGNIFICENT HUMMINGBIRD (Eugenes fulgens)
FIERY-THROATED HUMMINGBIRD (Panterpe insignis)
WHITE-BELLIED MOUNTAIN-GEM (Lampornis hemileucus)
PURPLE-THROATED MOUNTAIN-GEM (Lampornis calolaemus)
MAGENTA-THROATED WOODSTAR (Calliphlox bryantae)

Sungrebe was one of the big highlights of our boat trips at Caño Negro. Photo by guide Tom Johnson.
RUBY-THROATED HUMMINGBIRD (Archilochus colubris)
VIOLET-HEADED HUMMINGBIRD (Klais guimeti)
SCALY-BREASTED HUMMINGBIRD (Phaeochroa cuvierii)
VIOLET SABREWING (Campylopterus hemileucurus)
STRIPE-TAILED HUMMINGBIRD (Eupherusa eximia)
BLACK-BELLIED HUMMINGBIRD (Eupherusa nigriventris)
COPPERY-HEADED EMERALD (Elvira cupreiceps) [E]
RUFOUS-TAILED HUMMINGBIRD (Amazilia tzacatl)
Trogonidae (Trogons)
SLATY-TAILED TROGON (Trogon massena)
BLACK-HEADED TROGON (Trogon melanocephalus)

Spangle-cheeked Tanagers were regular visitors to the feeders at Bosque de Paz. Photo by guide Tom Johnson.
BLACK-THROATED TROGON (Trogon rufus)
ORANGE-BELLIED TROGON (Trogon aurantiiventris)
COLLARED TROGON (Trogon collaris)
Momotidae (Motmots)
KEEL-BILLED MOTMOT (Electron carinatum)
BROAD-BILLED MOTMOT (Electron platyrhynchum)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)
BELTED KINGFISHER (Megaceryle alcyon)
AMAZON KINGFISHER (Chloroceryle amazona)

Much to our amazement, White-throated Crakes paraded out in the open near La Fortuna. Photo by guide Tom Johnson.
GREEN KINGFISHER (Chloroceryle americana)
AMERICAN PYGMY KINGFISHER (Chloroceryle aenea)
Bucconidae (Puffbirds)
PIED PUFFBIRD (Notharchus tectus)
Galbulidae (Jacamars)
RUFOUS-TAILED JACAMAR (Galbula ruficauda)
Capitonidae (New World Barbets)
RED-HEADED BARBET (Eubucco bourcierii)
Semnornithidae (Toucan-Barbets)
PRONG-BILLED BARBET (Semnornis frantzii)
Ramphastidae (Toucans)
EMERALD TOUCANET (BLUE-THROATED) (Aulacorhynchus prasinus caeruleogularis)
COLLARED ARACARI (Pteroglossus torquatus)
YELLOW-EARED TOUCANET (Selenidera spectabilis)

We had some great luck with fruiting fig trees on this tour. Here, a Yellow-throated Toucan searches for its next morsel. Photo by guide Tom Johnson.
YELLOW-THROATED TOUCAN (CHESTNUT-MANDIBLED) (Ramphastos ambiguus swainsonii)
KEEL-BILLED TOUCAN (Ramphastos sulfuratus)
Picidae (Woodpeckers)
BLACK-CHEEKED WOODPECKER (Melanerpes pucherani)
HOFFMANN'S WOODPECKER (Melanerpes hoffmannii)
HAIRY WOODPECKER (Picoides villosus)
GOLDEN-OLIVE WOODPECKER (Colaptes rubiginosus)
LINEATED WOODPECKER (Dryocopus lineatus)
Falconidae (Falcons and Caracaras)
BARRED FOREST-FALCON (Micrastur ruficollis) [*]

An anteater with a pattern like a skunk! Northern Tamandua put on a good show for us along the Peninsula Road at Arenal, climbing down from a tree and then up a bank across a gully from us. Photo by guide Tom Johnson.
CRESTED CARACARA (Caracara cheriway)
LAUGHING FALCON (Herpetotheres cachinnans)
MERLIN (Falco columbarius)
BAT FALCON (Falco rufigularis)
Psittacidae (New World and African Parrots)
ORANGE-CHINNED PARAKEET (Brotogeris jugularis)
BROWN-HOODED PARROT (Pyrilia haematotis)
WHITE-CROWNED PARROT (Pionus senilis)
RED-LORED PARROT (Amazona autumnalis)
WHITE-FRONTED PARROT (Amazona albifrons)
OLIVE-THROATED PARAKEET (Eupsittula nana)
CRIMSON-FRONTED PARAKEET (Psittacara finschi)
Thamnophilidae (Typical Antbirds)
GREAT ANTSHRIKE (Taraba major)
BARRED ANTSHRIKE (Thamnophilus doliatus)

Violet Sabrewings often buzzed by just inches away near the feeders at Bosque de Paz. Photo by guide Cory Gregory.
RUSSET ANTSHRIKE (Thamnistes anabatinus)
STREAK-CROWNED ANTVIREO (Dysithamnus striaticeps)
SLATY ANTWREN (Myrmotherula schisticolor)
DUSKY ANTBIRD (Cercomacroides tyrannina)
BARE-CROWNED ANTBIRD (Gymnocichla nudiceps)
CHESTNUT-BACKED ANTBIRD (Poliocrania exsul)
DULL-MANTLED ANTBIRD (Sipia laemosticta)
SPOTTED ANTBIRD (Hylophylax naevioides)
OCELLATED ANTBIRD (Phaenostictus mcleannani) [*]
Grallariidae (Antpittas)
THICKET ANTPITTA (Hylopezus dives)
Furnariidae (Ovenbirds and Woodcreepers)
WEDGE-BILLED WOODCREEPER (Glyphorynchus spirurus)
COCOA WOODCREEPER (Xiphorhynchus susurrans)
SPOTTED WOODCREEPER (Xiphorhynchus erythropygius)

This Slate-colored Grosbeak called out right overhead at the Arenal waterfall trail. Photo by guide Tom Johnson.
BROWN-BILLED SCYTHEBILL (Campylorhamphus pusillus)
STREAK-HEADED WOODCREEPER (Lepidocolaptes souleyetii)
SPOT-CROWNED WOODCREEPER (Lepidocolaptes affinis)
SPOTTED BARBTAIL (Premnoplex brunnescens)
RUDDY TREERUNNER (Margarornis rubiginosus)
RED-FACED SPINETAIL (Cranioleuca erythrops)
Tyrannidae (Tyrant Flycatchers)
YELLOW-BELLIED/ BROWN-CAPPED TYRANNULET (Ornithion sp.)
NORTHERN BEARDLESS-TYRANNULET (Camptostoma imberbe)
YELLOW TYRANNULET (Capsiempis flaveola)
YELLOW-BELLIED ELAENIA (Elaenia flavogaster)
MOUNTAIN ELAENIA (Elaenia frantzii)

A pair of Streak-crowned Antvireos showed well along the waterfall trail at Arenal Observatory Lodge. Photo by guide Tom Johnson.
TORRENT TYRANNULET (Serpophaga cinerea)
OLIVE-STRIPED FLYCATCHER (Mionectes olivaceus)
PALTRY TYRANNULET (Zimmerius vilissimus)
SCALE-CRESTED PYGMY-TYRANT (Lophotriccus pileatus)
COMMON TODY-FLYCATCHER (Todirostrum cinereum)
BLACK-HEADED TODY-FLYCATCHER (Todirostrum nigriceps)
TUFTED FLYCATCHER (Mitrephanes phaeocercus)
DARK PEWEE (Contopus lugubris)
TROPICAL PEWEE (Contopus cinereus)
YELLOW-BELLIED FLYCATCHER (Empidonax flaviventris)

Our close observations of Prong-billed Barbets allowed us to see the bristles at the base of the bill and the prongs of the bill, too. Photo by guide Tom Johnson.
ACADIAN FLYCATCHER (Empidonax virescens)
YELLOWISH FLYCATCHER (Empidonax flavescens)
BLACK PHOEBE (Sayornis nigricans)
LONG-TAILED TYRANT (Colonia colonus)
BRIGHT-RUMPED ATTILA (Attila spadiceus)
DUSKY-CAPPED FLYCATCHER (Myiarchus tuberculifer)
GREAT CRESTED FLYCATCHER (Myiarchus crinitus)
GREAT KISKADEE (Pitangus sulphuratus)
BOAT-BILLED FLYCATCHER (Megarynchus pitangua)
SOCIAL FLYCATCHER (Myiozetetes similis)

This lovely Scaled Pigeon flew in and perched along the road west of Caño Negro, allowing guide Cory Gregory to snap this great digiscoped shot. Check out that awesome scaling!
GRAY-CAPPED FLYCATCHER (Myiozetetes granadensis)
TROPICAL KINGBIRD (Tyrannus melancholicus)
FORK-TAILED FLYCATCHER (Tyrannus savana)
Pipridae (Manakins)
WHITE-RUFFED MANAKIN (Corapipo altera)
WHITE-COLLARED MANAKIN (Manacus candei)
Tityridae (Tityras and Allies)
BLACK-CROWNED TITYRA (Tityra inquisitor)
MASKED TITYRA (Tityra semifasciata)
CINNAMON BECARD (Pachyramphus cinnamomeus)
Vireonidae (Vireos, Shrike-Babblers, and Erpornis)
LESSER GREENLET (Pachysylvia decurtata)

This Paca was one of several that visited us at night at Bosque de Paz. Photo by guide Tom Johnson.
YELLOW-THROATED VIREO (Vireo flavifrons)
BROWN-CAPPED VIREO (Vireo leucophrys)
Corvidae (Crows, Jays, and Magpies)
AZURE-HOODED JAY (Cyanolyca cucullata)
BROWN JAY (Psilorhinus morio)
Hirundinidae (Swallows)
BLUE-AND-WHITE SWALLOW (Pygochelidon cyanoleuca)
NORTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx serripennis)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)
GRAY-BREASTED MARTIN (Progne chalybea)
MANGROVE SWALLOW (Tachycineta albilinea)
Troglodytidae (Wrens)
NIGHTINGALE WREN (Microcerculus philomela) [*]
HOUSE WREN (Troglodytes aedon)
OCHRACEOUS WREN (Troglodytes ochraceus)

A Hepatic Tanager female dropped down to the ground to pursue this fancy mantis, who was not too pleased with the encounter. Photo by guide Cory Gregory.
BAND-BACKED WREN (Campylorhynchus zonatus)
SPOT-BREASTED WREN (Pheugopedius maculipectus)
BLACK-THROATED WREN (Pheugopedius atrogularis)
STRIPE-BREASTED WREN (Cantorchilus thoracicus)
CABANIS'S WREN (Cantorchilus modestus)
CANEBRAKE WREN (Cantorchilus zeledoni)

The fig trees at Arenal attracted a wide variety of neat songbirds including this dramatic-looking Masked Tityra. Photo by guide Tom Johnson.
BAY WREN (Cantorchilus nigricapillus)
WHITE-BREASTED WOOD-WREN (Henicorhina leucosticta)
GRAY-BREASTED WOOD-WREN (Henicorhina leucophrys)
SONG WREN (Cyphorhinus phaeocephalus)
Polioptilidae (Gnatcatchers)
TAWNY-FACED GNATWREN (Microbates cinereiventris)
Cinclidae (Dippers)
AMERICAN DIPPER (Cinclus mexicanus)
Turdidae (Thrushes and Allies)
BLACK-FACED SOLITAIRE (Myadestes melanops)
BLACK-BILLED NIGHTINGALE-THRUSH (Catharus gracilirostris)
SLATY-BACKED NIGHTINGALE-THRUSH (Catharus fuscater)
RUDDY-CAPPED NIGHTINGALE-THRUSH (Catharus frantzii)

Our boat trips at Medio Queso and Caño Negro were great for seeing Mangrove Cuckoos this year. Photo by guide Tom Johnson.
BLACK-HEADED NIGHTINGALE-THRUSH (Catharus mexicanus)
WOOD THRUSH (Hylocichla mustelina)
SOOTY THRUSH (Turdus nigrescens)
MOUNTAIN THRUSH (Turdus plebejus)
CLAY-COLORED THRUSH (Turdus grayi)
WHITE-THROATED THRUSH (Turdus assimilis)
Mimidae (Mockingbirds and Thrashers)
GRAY CATBIRD (Dumetella carolinensis)
TROPICAL MOCKINGBIRD (Mimus gilvus)
Ptiliogonatidae (Silky-flycatchers)
BLACK-AND-YELLOW SILKY-FLYCATCHER (Phainoptila melanoxantha)

This Coppery-headed Emerald was a fancy endemic species that we saw really well at the La Paz Waterfall Gardens. Photo by guide Cory Gregory.
LONG-TAILED SILKY-FLYCATCHER (Ptiliogonys caudatus)
Parulidae (New World Warblers)
LOUISIANA WATERTHRUSH (Parkesia motacilla)
NORTHERN WATERTHRUSH (Parkesia noveboracensis)
GOLDEN-WINGED WARBLER (Vermivora chrysoptera)
BLACK-AND-WHITE WARBLER (Mniotilta varia)
PROTHONOTARY WARBLER (Protonotaria citrea)
FLAME-THROATED WARBLER (Oreothlypis gutturalis)
TENNESSEE WARBLER (Oreothlypis peregrina)
GRAY-CROWNED YELLOWTHROAT (Geothlypis poliocephala)
MOURNING WARBLER (Geothlypis philadelphia)
KENTUCKY WARBLER (Geothlypis formosa)
OLIVE-CROWNED YELLOWTHROAT (Geothlypis semiflava)

These Black-bellied Whistling-Ducks were part of a large flock at the Medio Queso wetlands. Photo by guide Cory Gregory.
COMMON YELLOWTHROAT (Geothlypis trichas)
TROPICAL PARULA (Setophaga pitiayumi)
YELLOW WARBLER (Setophaga petechia)
CHESTNUT-SIDED WARBLER (Setophaga pensylvanica)
BLACK-THROATED GREEN WARBLER (Setophaga virens)
BLACK-CHEEKED WARBLER (Basileuterus melanogenys)
GOLDEN-CROWNED WARBLER (Basileuterus culicivorus)
COSTA RICAN WARBLER (Basileuterus melanotis)
BUFF-RUMPED WARBLER (Myiothlypis fulvicauda)

An Olive-crowned Yellowthroat took a break from singing to check out our boat on the Caño Negro. Photo by guide Tom Johnson.
WILSON'S WARBLER (Cardellina pusilla)
SLATE-THROATED REDSTART (Myioborus miniatus)
Thraupidae (Tanagers and Allies)
WHITE-THROATED SHRIKE-TANAGER (Lanio leucothorax)
CRIMSON-COLLARED TANAGER (Ramphocelus sanguinolentus)
PASSERINI'S TANAGER (Ramphocelus passerinii)
BLUE-GRAY TANAGER (Thraupis episcopus)
PALM TANAGER (Thraupis palmarum)
GOLDEN-HOODED TANAGER (Tangara larvata)
SPANGLE-CHEEKED TANAGER (Tangara dowii)
RUFOUS-WINGED TANAGER (Tangara lavinia)

One of our best sightings on the Caño Negro boat trips was this Gray-headed Kite that perched up beautifully along the riverbank. Photo by guide Tom Johnson.
BAY-HEADED TANAGER (Tangara gyrola)
EMERALD TANAGER (Tangara florida)
SILVER-THROATED TANAGER (Tangara icterocephala)
RED-LEGGED HONEYCREEPER (Cyanerpes cyaneus)
GREEN HONEYCREEPER (Chlorophanes spiza)
SLATY FLOWERPIERCER (Diglossa plumbea)
BLUE-BLACK GRASSQUIT (Volatinia jacarina)
THICK-BILLED SEED-FINCH (Sporophila funerea)
NICARAGUAN SEED-FINCH (Sporophila nuttingi)
VARIABLE SEEDEATER (Sporophila corvina)
WHITE-COLLARED SEEDEATER (Sporophila torqueola)
BANANAQUIT (Coereba flaveola)
YELLOW-FACED GRASSQUIT (Tiaris olivaceus)
BUFF-THROATED SALTATOR (Saltator maximus)

Our time with Golden-browed Chlorophonias on this trip was magical! Look at that combination of colors! Photo by guide Tom Johnson.
GRAYISH SALTATOR (Saltator coerulescens)
SLATE-COLORED GROSBEAK (Saltator grossus)
Emberizidae (Buntings and New World Sparrows)
SOOTY-CAPPED CHLOROSPINGUS (Chlorospingus pileatus)
COMMON CHLOROSPINGUS (Chlorospingus flavopectus)
BLACK-STRIPED SPARROW (Arremonops conirostris)
ORANGE-BILLED SPARROW (Arremon aurantiirostris)
CHESTNUT-CAPPED BRUSHFINCH (Arremon brunneinucha)
SOOTY-FACED FINCH (Arremon crassirostris)
RUFOUS-COLLARED SPARROW (Zonotrichia capensis)
YELLOW-THIGHED FINCH (Pselliophorus tibialis)
Cardinalidae (Cardinals and Allies)
HEPATIC TANAGER (Piranga flava)

This tiny pescador, the American Pygmy-Kingfisher, was hunting along the edge of the water at Medio Queso. Photo by guide Tom Johnson.
SUMMER TANAGER (Piranga rubra)
CARMIOL'S TANAGER (Chlorothraupis carmioli)
Icteridae (Troupials and Allies)
RED-WINGED BLACKBIRD (Agelaius phoeniceus)
RED-BREASTED MEADOWLARK (Sturnella militaris)
MELODIOUS BLACKBIRD (Dives dives)
GREAT-TAILED GRACKLE (Quiscalus mexicanus)
NICARAGUAN GRACKLE (Quiscalus nicaraguensis)
BRONZED COWBIRD (Molothrus aeneus)
BLACK-COWLED ORIOLE (Icterus prosthemelas)
ORCHARD ORIOLE (Icterus spurius)
BALTIMORE ORIOLE (Icterus galbula)
MONTEZUMA OROPENDOLA (Psarocolius montezuma)
Fringillidae (Finches, Euphonias, and Allies)
YELLOW-THROATED EUPHONIA (Euphonia hirundinacea)
OLIVE-BACKED EUPHONIA (Euphonia gouldi)

Emerald Tanager was a regular member of the flock feeding on fruit in the garden at Arenal Observatory Lodge. Photo by guide Tom Johnson.
TAWNY-CAPPED EUPHONIA (Euphonia anneae)
GOLDEN-BROWED CHLOROPHONIA (Chlorophonia callophrys)
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus) [I]
COMMON OPOSSUM (Didelphis marsupialis)
MANTLED HOWLER MONKEY (Alouatta palliata)
WHITE-THROATED CAPUCHIN (Cebus capucinus)
CENTRAL AMERICAN SPIDER MONKEY (Ateles geoffroyi)
NORTHERN TAMANDUA (Tamandua mexicana)
VARIEGATED SQUIRREL (Sciurus variegatoides)
RED-TAILED SQUIRREL (Sciurus granatensis)
CENTRAL AMERICAN AGOUTI (Dasyprocta punctata)
PACA (Cuniculus paca)

Pinnated Bitterns showed very well along the edge of the river at Caño Negro. We were pretty fortunate to see this sneaky heron this well. Photo by guide Tom Johnson.
NORTHERN RACCOON (Procyon lotor)
WHITE-NOSED COATI (Nasua narica)
Herps
GREEN IGUANA (Iguana iguana)
TROPICAL HOUSE GECKO (Hemidactylus mabouia)
CENTRAL AMERICAN WHIPTAIL (Ameiva festiva)
SPECTACLED CAIMAN (Caiman crocodilus)
CANE TOAD (Rhinella marina)
Totals for the tour: 306 bird taxa and 11 mammal taxa