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Early morning sunlight bathes the Parthenon and Acropolis, as seen from the rooftop restaurant of our nearby hotel. Photo by guide Megan Edwards Crewe.
Greece's lovely countryside offers a delightful place to observe Europe's spring migration and to study some of its breeding birds -- with the added bonus of a plethora of nearby cultural attractions to enjoy as well! Our two-week tour ventured from the wild pine forests of Dadia NP in the country's northeast to teeming wetlands along the calm Aegean Sea, from the fertile countryside around the massive, freshwater Lake Kerkini to the brooding hulks of Mount Olympus and Mount Parnassos, and the dry, olive-studded hillsides around Delphi. Our weather was generally delightful (except for those two dampish days at the beginning, of course), and we had a multitude of birds to enjoy.
At Dadia's raptor feeding station, an aggressive scrum around the carcasses held dozens of Cinereous Vultures and Eurasian Griffons, with a few Egyptian Vultures, a single White-tailed Eagle and some Black Kites watching the action from a nearby tree. Great Crested Grebes performed their courtship mirror dances and gathered huge mouthfuls of marsh vegetation to add to their growing platform nests. The lovely songs of a bevy of larks -- Calandra, Wood, Greater Short-toed, Crested, Sky -- rained down from the heavens as the birds themselves hovered in endless display flights. A White Stork strode through a field studded with blood-red poppies (and scores more snoozed on nests all across northern Greece). A pale pink cloud of Greater Flamingos floated on long legs along the edge of a lagoon. A Syrian Woodpecker hitched its way up a telephone pole. Sardinian and Subalpine and Rueppell's and Olivaceous warblers sang from nearby bush tops, conveniently right at eye level mere yards away, while Olive-tree Warblers led us on quite the merry dance before one FINALLY showed itself -- appropriately enough in a little olive tree. Masked Shrikes hunted from low branches around our picnic site. A Little Owl sat on a hay bale, staring intently at something on the ground. Dalmatian Pelicans preened on a muddy islet, looking especially enormous compared to nearby Pygmy Cormorants. Two Rock Nuthatches ferried mouthful after mouthful of insects to a nest stuck to the wall of the Athenian Treasury at Delphi. Short-toed Eagles hovered over farm fields. Common Cuckoos shouted challenges from wooded hillsides. Red-footed Falcons rested on telephone wires. Spur-winged Plovers trotted across dry pans. Alpine Swifts raced in screaming groups over a craggy headland, with the blue, blue sea behind them. A Firecrest flared his stunning crown feathers. A pair of Hawfinches rummaged along a mountain roadside. And technicolor European Bee-eaters showed repeatedly that not all European birds are boring little brown things!
Of course, it wasn't just the birds that made this trip. The Parthenon and Acropolis were still impressive, even though clad in scaffolding, and the new museum there is fabulous. In Iasmos gorge, we walked on the graceful spans of an old stone bridge that dates back to Byzantine times. The grave treasures of Philip II (former king of Macedonia and father of Alexander the Great) at Vergina dazzled in their spectacular museum. The ancient ruins at Delphi, once thought to be the center of the universe, sprawl down the dry, maquis-studded slopes, the writings on their many stone blocks still perfectly legible (and readable, if you can read Greek) millennia later. And a quick visit to the glittering mosaics of Osios Loukas finished off our tour.
And of course, the trip was greatly enhanced by having such a congenial and enjoyable bunch of participants to share it with. Thanks so much to all of you for joining me for the adventure. Thanks too to Lefteris -- for both his translation skills and his excellent bird-spotting! Thanks to Eleni, Christina and Irini for their excellent ruins tours, and to Sharon at FG headquarters for putting it all together. I hope to see you all again in the field somewhere soon!
-- Megan
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant

A Great Crested Grebe, in all its breeding finery, floats on Lake Kerkini. We saw hundreds dotted across the surface of the water, including many doing their endearing courtship mirror dances or building their soggy platform nests. Photo by participant Merrill Lester.
Anatidae (Ducks, Geese, and Waterfowl)
GRAYLAG GOOSE (Anser anser)
MUTE SWAN (Cygnus olor)
COMMON SHELDUCK (Tadorna tadorna)
MALLARD (Anas platyrhynchos)
NORTHERN SHOVELER (Anas clypeata)
NORTHERN PINTAIL (Anas acuta)
GARGANEY (Anas querquedula)
FERRUGINOUS DUCK (Aythya nyroca)

A male Sardinian Warbler sang his challenges from the top of a tree near the edge of the Athens Agora. Photo by participant Merrill Lester.
Phasianidae (Pheasants, Grouse, and Allies)
RING-NECKED PHEASANT (Phasianus colchicus) [I*]
Gaviidae (Loons)
ARCTIC LOON (Gavia arctica)
Podicipedidae (Grebes)
LITTLE GREBE (Tachybaptus ruficollis)
GREAT CRESTED GREBE (Podiceps cristatus) [N]
Phoenicopteridae (Flamingos)
GREATER FLAMINGO (Phoenicopterus roseus)
Procellariidae (Shearwaters and Petrels)
YELKOUAN SHEARWATER (Puffinus yelkouan)
Ciconiidae (Storks)
BLACK STORK (Ciconia nigra)
WHITE STORK (Ciconia ciconia) [N]
Phalacrocoracidae (Cormorants and Shags)
GREAT CORMORANT (Phalacrocorax carbo) [N]
EUROPEAN SHAG (Phalacrocorax aristotelis)
PYGMY CORMORANT (Phalacrocorax pygmeus)
Pelecanidae (Pelicans)
GREAT WHITE PELICAN (Pelecanus onocrotalus)
DALMATIAN PELICAN (Pelecanus crispus)

A Short-toed Snake-Eagle on the wing, searching for its next meal. The copious covering of feathers on its thick, puffy head helps to protect it from snake strikes. Photo by participant Merrill Lester.
Ardeidae (Herons, Egrets, and Bitterns)
LITTLE BITTERN (Ixobrychus minutus)
GRAY HERON (Ardea cinerea)
PURPLE HERON (Ardea purpurea)
GREAT EGRET (Ardea alba)
LITTLE EGRET (Egretta garzetta)
CATTLE EGRET (Bubulcus ibis)
SQUACCO HERON (Ardeola ralloides)
BLACK-CROWNED NIGHT-HERON (Nycticorax nycticorax)

Little Owls were common and widespread this year, with at least a half-dozen sightings. Photo by participant Merrill Lester.
Threskiornithidae (Ibises and Spoonbills)
GLOSSY IBIS (Plegadis falcinellus)
EURASIAN SPOONBILL (Platalea leucorodia)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus)
Accipitridae (Hawks, Eagles, and Kites)
EGYPTIAN VULTURE (Neophron percnopterus)
EUROPEAN HONEY-BUZZARD (Pernis apivorus)
CINEREOUS VULTURE (Aegypius monachus)
EURASIAN GRIFFON (Gyps fulvus)
SHORT-TOED SNAKE-EAGLE (Circaetus gallicus)
LESSER SPOTTED EAGLE (Clanga pomarina)
BOOTED EAGLE (Hieraaetus pennatus)
EURASIAN MARSH-HARRIER (Circus aeruginosus)
LEVANT SPARROWHAWK (Accipiter brevipes)
EURASIAN SPARROWHAWK (Accipiter nisus)

A Gray Heron takes flight over a marshy edge of Lake Kerkini. This man-made reservoir has quickly become an important stopover spot for migrant waterfowl, shorebirds, egrets, herons, and more -- as well as a breeding area for pelicans and cormorants. Photo by participant Merrill Lester.
BLACK KITE (Milvus migrans)
WHITE-TAILED EAGLE (Haliaeetus albicilla)
COMMON BUZZARD (Buteo buteo)
Rallidae (Rails, Gallinules, and Coots)
EURASIAN MOORHEN (Gallinula chloropus)
EURASIAN COOT (Fulica atra)
Burhinidae (Thick-knees)
EURASIAN THICK-KNEE (Burhinus oedicnemus)

European Stonechats thrive on the dry, sunny, maquis-covered slopes around Delphi. Photo by participant Merrill Lester.
Recurvirostridae (Stilts and Avocets)
BLACK-WINGED STILT (Himantopus himantopus)
PIED AVOCET (Recurvirostra avosetta)
Haematopodidae (Oystercatchers)
EURASIAN OYSTERCATCHER (Haematopus ostralegus)
Charadriidae (Plovers and Lapwings)
BLACK-BELLIED PLOVER (Pluvialis squatarola)
SPUR-WINGED LAPWING (Vanellus spinosus)
KENTISH PLOVER (KENTISH) (Charadrius alexandrinus alexandrinus)
COMMON RINGED PLOVER (Charadrius hiaticula)
LITTLE RINGED PLOVER (Charadrius dubius)
Scolopacidae (Sandpipers and Allies)
SPOTTED REDSHANK (Tringa erythropus)
COMMON GREENSHANK (Tringa nebularia)
MARSH SANDPIPER (Tringa stagnatilis)
WOOD SANDPIPER (Tringa glareola)
COMMON REDSHANK (Tringa totanus)
EURASIAN CURLEW (Numenius arquata)

A scrum of Cinereous Vultures and Eurasian Griffons surround the carcass piles at the Dadia Raptor Center's feeding station, while an Egyptian Vulture, a Red Kite, an immature White-tailed Eagle and more Cinereous Vultures look on from an overhanging tree. Photo by guide Megan Edwards Crewe.
RUDDY TURNSTONE (Arenaria interpres)
CURLEW SANDPIPER (Calidris ferruginea)
DUNLIN (Calidris alpina)
LITTLE STINT (Calidris minuta)
Glareolidae (Pratincoles and Coursers)
COLLARED PRATINCOLE (Glareola pratincola)
Laridae (Gulls, Terns, and Skimmers)
SLENDER-BILLED GULL (Chroicocephalus genei)

Finding a Great Spotted Woodpecker nest allowed us some fine, close observations of the busy parents. Photo by participant Merrill Lester.
BLACK-HEADED GULL (Chroicocephalus ridibundus)
MEDITERRANEAN GULL (Ichthyaetus melanocephalus)
YELLOW-LEGGED GULL (MICHAHELLIS) (Larus michahellis michahellis)
LITTLE TERN (Sternula albifrons)
GULL-BILLED TERN (Gelochelidon nilotica)
BLACK TERN (Chlidonias niger)
WHITE-WINGED TERN (Chlidonias leucopterus)
WHISKERED TERN (Chlidonias hybrida)
COMMON TERN (Sterna hirundo) [N]
SANDWICH TERN (Thalasseus sandvicensis)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia)
COMMON WOOD-PIGEON (Columba palumbus)
EUROPEAN TURTLE-DOVE (Streptopelia turtur)
EURASIAN COLLARED-DOVE (Streptopelia decaocto)
Cuculidae (Cuckoos)
COMMON CUCKOO (Cuculus canorus)
Strigidae (Owls)
LITTLE OWL (Athene noctua)

The elegant Erechtheion is a smaller temple that flanks the Parthenon atop the Acropolis in Athens. The six caryatids (pillars carved to look like women) are replicas; the originals have been moved to the fabulous new Parthenon museum. Photo by guide Megan Edwards Crewe.
Apodidae (Swifts)
ALPINE SWIFT (Apus melba)
COMMON SWIFT (Apus apus)
PALLID SWIFT (Apus pallidus)
Upupidae (Hoopoes)
EURASIAN HOOPOE (Upupa epops) [N]
Alcedinidae (Kingfishers)
COMMON KINGFISHER (Alcedo atthis)

In Greece, the charming little European Robin is restricted to higher elevations -- like the slopes of Mount Olympus, where this one was photographed. Photo by participant Merrill Lester.
Meropidae (Bee-eaters)
EUROPEAN BEE-EATER (Merops apiaster) [N]
Coraciidae (Rollers)
EUROPEAN ROLLER (Coracias garrulus)
Picidae (Woodpeckers)
EURASIAN WRYNECK (Jynx torquilla)
GREAT SPOTTED WOODPECKER (Dendrocopos major) [N]
SYRIAN WOODPECKER (Dendrocopos syriacus)
BLACK WOODPECKER (Dryocopus martius)
EURASIAN GREEN WOODPECKER (Picus viridis) [*]
Falconidae (Falcons and Caracaras)
LESSER KESTREL (Falco naumanni)
EURASIAN KESTREL (Falco tinnunculus) [N]
RED-FOOTED FALCON (Falco vespertinus)
EURASIAN HOBBY (Falco subbuteo)
PEREGRINE FALCON (Falco peregrinus)

A Yellow-legged Gull keeps an eye on things from its rooftop perch at Agios Nikolaos (Saint Nicholas's church) near Porto Lagos. Photo by participant Merrill Lester.
Laniidae (Shrikes)
RED-BACKED SHRIKE (Lanius collurio)
LESSER GRAY SHRIKE (Lanius minor)
MASKED SHRIKE (Lanius nubicus)
WOODCHAT SHRIKE (Lanius senator)
Oriolidae (Old World Orioles)
EURASIAN GOLDEN ORIOLE (Oriolus oriolus)
Corvidae (Crows, Jays, and Magpies)
EURASIAN JAY (Garrulus glandarius)

The loud, cheery songs of the Common Chaffinch were a regular part of the tour's soundtrack -- particularly in Dadia and around Delphi. Photo by participant Merrill Lester.
EURASIAN MAGPIE (Pica pica)
YELLOW-BILLED CHOUGH (Pyrrhocorax graculus)
EURASIAN JACKDAW (Corvus monedula) [N]
HOODED CROW (Corvus cornix)
COMMON RAVEN (Corvus corax)
Alaudidae (Larks)
CALANDRA LARK (Melanocorypha calandra)
GREATER SHORT-TOED LARK (Calandrella brachydactyla) [*]
CRESTED LARK (Galerida cristata)
SKY LARK (Alauda arvensis)
WOOD LARK (Lullula arborea)
Hirundinidae (Swallows)
BANK SWALLOW (Riparia riparia)
EURASIAN CRAG-MARTIN (Ptyonoprogne rupestris)
BARN SWALLOW (Hirundo rustica) [N]
RED-RUMPED SWALLOW (Cecropis daurica) [N]
COMMON HOUSE-MARTIN (Delichon urbicum) [N]
A White Stork forages in a lovely field full of wildflowers near Lake Kerkini while a host of frogs and breeding birds shouts in the background. Video by guide Megan Edwards Crewe.
Paridae (Tits, Chickadees, and Titmice)
COAL TIT (Periparus ater)
CRESTED TIT (Lophophanes cristatus)
SOMBRE TIT (Poecile lugubris)
EURASIAN BLUE TIT (Cyanistes caeruleus)
GREAT TIT (Parus major)
Remizidae (Penduline-Tits)
EURASIAN PENDULINE-TIT (Remiz pendulinus)
Aegithalidae (Long-tailed Tits)
LONG-TAILED TIT (Aegithalos caudatus)
Sittidae (Nuthatches)
EURASIAN NUTHATCH (Sitta europaea) [N]

A male Subalpine Warbler pauses for a moment, resting between trips to its nest with mouthfuls of food. Photo by participant Merrill Lester.
ROCK NUTHATCH (Sitta neumayer) [N]
Certhiidae (Treecreepers)
SHORT-TOED TREECREEPER (Certhia brachydactyla)
Troglodytidae (Wrens)
EURASIAN WREN (Troglodytes troglodytes)
Regulidae (Kinglets)
FIRECREST (Regulus ignicapilla)
Cettiidae (Bush-Warblers and Allies)
CETTI'S WARBLER (Cettia cetti)

The wildflowers were blooming all over Greece, making for some lovely backdrops. This is a clover, but I'm not sure which one. Photo by guide Megan Edwards Crewe.
Phylloscopidae (Leaf Warblers)
COMMON CHIFFCHAFF (Phylloscopus collybita) [*]
EASTERN BONELLI'S WARBLER (Phylloscopus orientalis)
Acrocephalidae (Reed-Warblers and Allies)
EASTERN OLIVACEOUS WARBLER (Iduna pallida)
OLIVE-TREE WARBLER (Hippolais olivetorum)
EURASIAN REED-WARBLER (Acrocephalus scirpaceus)
GREAT REED-WARBLER (Acrocephalus arundinaceus)
Cisticolidae (Cisticolas and Allies)
ZITTING CISTICOLA (Cisticola juncidis)
Sylviidae (Sylviid Warblers)
EURASIAN BLACKCAP (Sylvia atricapilla)
LESSER WHITETHROAT (Sylvia curruca)
EASTERN ORPHEAN WARBLER (Sylvia crassirostris)
RUEPPELL'S WARBLER (Sylvia rueppelli)
SUBALPINE WARBLER (Sylvia cantillans)
SARDINIAN WARBLER (Sylvia melanocephala)
GREATER WHITETHROAT (Sylvia communis)

The gang takes a break for a picture along the edge of Lake Kerkini. The mountains in the background form the country's border with Bulgaria. Photo by guide Megan Edwards Crewe.
Muscicapidae (Old World Flycatchers)
SPOTTED FLYCATCHER (Muscicapa striata) [N]
EUROPEAN ROBIN (Erithacus rubecula)
COMMON NIGHTINGALE (Luscinia megarhynchos)
EUROPEAN PIED FLYCATCHER (Ficedula hypoleuca)
BLACK REDSTART (Phoenicurus ochruros)
RUFOUS-TAILED ROCK-THRUSH (Monticola saxatilis)
BLUE ROCK-THRUSH (Monticola solitarius)

Rock Nuthatches were abundant around Delphi; this one was nesting right on the Athenian Treasury building! Photo by participant Merrill Lester.
EUROPEAN STONECHAT (Saxicola rubicola rubicola)
NORTHERN WHEATEAR (Oenanthe oenanthe)
BLACK-EARED WHEATEAR (Oenanthe hispanica)
Turdidae (Thrushes and Allies)
EURASIAN BLACKBIRD (Turdus merula)
MISTLE THRUSH (Turdus viscivorus) [*]
Sturnidae (Starlings)
EUROPEAN STARLING (Sturnus vulgaris)
Motacillidae (Wagtails and Pipits)
WESTERN YELLOW WAGTAIL (BLACK-HEADED) (Motacilla flava feldegg)
GRAY WAGTAIL (Motacilla cinerea)
WHITE WAGTAIL (WHITE-FACED) (Motacilla alba alba)
TAWNY PIPIT (Anthus campestris)
Emberizidae (Buntings and New World Sparrows)
CIRL BUNTING (Emberiza cirlus)
ORTOLAN BUNTING (Emberiza hortulana)
BLACK-HEADED BUNTING (Emberiza melanocephala)
CORN BUNTING (Emberiza calandra)
Fringillidae (Finches, Euphonias, and Allies)
COMMON CHAFFINCH (Fringilla coelebs)

An Ortolan Bunting sings near Doriskos, unaware that drawing attention to himself might some day consign him to the pot. The French still regularly eat Ortolans as a delicacy, despite the fact that hunting and capturing them is now illegal. Photo by guide Megan Edwards Crewe.
EURASIAN BULLFINCH (Pyrrhula pyrrhula) [*]
EUROPEAN GREENFINCH (Chloris chloris aurantiiventris)
EUROPEAN GOLDFINCH (Carduelis carduelis)
EURASIAN LINNET (Carduelis cannabina)
EUROPEAN SERIN (Serinus serinus)
HAWFINCH (Coccothraustes coccothraustes) [N]
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus) [N]

Great Tits were widespread, found on many days of the tour. Photo by participant Merrill Lester.
SPANISH SPARROW (Passer hispaniolensis) [N]
EURASIAN TREE SPARROW (Passer montanus)
ROCK PETRONIA (Petronia petronia)
EUROPEAN RED SQUIRREL (Sciurus vulgaris)
NUTRIA (Myocastor coypus)
Herps
EUROPEAN POND TURTLE (Emys orbicularis)
MARGINATED TORTOISE (Testudo marginata)
BALKAN (GREEK MARSH) FROG (Pelophylax kurtmuelleri)
MARSH FROG (Pelophylax ridibundus)
EUROPEAN GREEN LIZARD (Lacerta viridis)
ERHARD'S WALL LIZARD (Podarcis erhardii)
Totals for the tour: 181 bird taxa and 2 mammal taxa