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The South Tufa at Mono Lake set an otherworldly scene for one stellar morning of birding east of the Sierra Nevada - from phalaropes and grebes to shrikes and sparrows. Photo by guide Tom Johnson.
Late August and early September mark a great point in the calendar to investigate a broad swath of biogeography in California, certainly one of the USA's birdiest states. Fall migration is in full swing for songbirds, the shorebird show is primed, and the seabird spectacle close to shore is, well, spectacular.
We began the tour with a nice sampling of habitats along the southwestern edge of San Francisco Bay, enjoying regional specialties like the newly split Ridgway's Rail and the spectacle of many thousands of shorebirds. Heading east across the Diablo Mountains, we visited several drier habitats. In the oak grasslands and scrubby slopes of Mines Rd. and Del Puerto Canyon, we found the California endemic Yellow-billed Magpie and the recently split Bell's Sparrow, along with the highly charismatic Lewis's Woodpecker and mysterious Lawrence's Goldfinch. We then continued east to the foot of the Sierra in Sonora, where we spent two nights.
Our first day of birding in the Sierra Nevada featured the Giant Sequoia grove at Calaveras Big Trees State Park, full of birds including a group favorite -White-headed Woodpecker. We also had amazingly close views of an enchanting Pacific Wren as it sang its complex, bubbly song from a brushpile. Heading higher into the mountains, we began our careful search for Mountain Quail, with a long stalk eventually culminating in good views of the quail for all members of the group. Amusingly, as soon as we started downhill after a long search on foot, Chris spotted a Mountain Quail standing on the side of the highway!
The next was spent birding our way over the Sierra crest at Sonora Pass en route to Lee Vining and Mono Lake. In the mountains, we enjoyed such montane specialties as American Dipper, Thick-billed Fox Sparrow, and Olive-sided Flycatcher. We even ran across some nice flocks of migrant songbirds, including many warblers and a super vocal Cassin's Vireo.
After an enjoyable time in the mountains, the high desert, conifer forest, and hypersaline lake habitat contrasts of the Mono Basin made for an otherworldly scene during our two night stay in Lee Vining. Highlights here included the legions of Red-necked Phalaropes, American Avocets, Eared Grebes, and California Gulls feeding against the backdrop of volcano cones, tall mountains, and stark limestone tufa formations in the storied waters of Mono Lake. Trips around the Mono Basin yielded such amazing birds as Greater Sage-Grouse, Black-backed Woodpecker, Williamson's Sapsucker, Pinyon Jay, Clark's Nutcracker, and Red Crossbill.
After an inspiring visit to the east side of the Sierra Nevada, we left the Mono Lake area and had a long day of driving back to the west, ending on the Pacific Ocean at Half Moon Bay where we stayed through until the end of the tour. This leg of the journey was anchored by a marvelous pelagic trip out of Half Moon Bay on the Huli Cat, organized by our friend Alvaro Jaramillo of Alvaro's Adventures. On relatively placid seas, we absorbed many encounters with a strong cast of seabirds, including six alcids and ten tubenoses. Highlights of the trip included two close Tufted Puffins in breeding plumage, a few close passes from a Flesh-footed Shearwater, and a close and stunning Laysan Albatross that was camped out alongside a fishing vessel along with almost 40 Black-footed Albatrosses. Photos of the Laysan Albie documented a set of leg bands, including a clear shot of its unique identifying code! Mammals were also great, with a duo of Dall's Porpoise (AKA Panda Tuna) streaking in from the port side to ride under the boat's bow.
A final day along the San Mateo Co. coast led us through Monterey Cypress clumps and rocky promontories along the ocean and up into a grove of impressive Coastal Redwoods in search of Wrentit, Townsend's Warbler, Snowy Plover, and Wandering Tattler. We even found a crisp juvenile White-tailed Kite and got to watch an offshore feeding frenzy of seabirds and Humpback Whales at Pescadero State Beach. A nice Italian dinner with a finishing touch of creme brulee was the perfect way to cap off this tour as the sun sank into the Pacific Ocean.
Happily, we ended up having a perfect run of weather, and a long list of excellent birds followed suit. Chris and I really enjoyed traveling with everyone through such a beautiful and diverse array of habitats. We both hope to see you out in the field again soon.
-Tom Johnson
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Anatidae (Ducks, Geese, and Waterfowl)
CANADA GOOSE (Branta canadensis)
GADWALL (Anas strepera)
MALLARD (Anas platyrhynchos)
CINNAMON TEAL (Anas cyanoptera)
NORTHERN SHOVELER (Anas clypeata)

Mountain Quail proved to be quite popular - after an initial struggle to find these skulky forest quail, we succeeded - and then had several more opportunities to see them alongside roads in the Sierra Nevada. Photo by guide Tom Johnson.
NORTHERN PINTAIL (Anas acuta) GREEN-WINGED TEAL (AMERICAN) (Anas crecca carolinensis)
HARLEQUIN DUCK (Histrionicus histrionicus)
SURF SCOTER (Melanitta perspicillata)
RUDDY DUCK (Oxyura jamaicensis)
Odontophoridae (New World Quail)
MOUNTAIN QUAIL (Oreortyx pictus)
CALIFORNIA QUAIL (Callipepla californica)
Phasianidae (Pheasants, Grouse, and Allies)
GREATER SAGE-GROUSE (Centrocercus urophasianus)
WILD TURKEY (Meleagris gallopavo)
Gaviidae (Loons)
RED-THROATED LOON (Gavia stellata)

This juvenile Laysan Albatross, marked by color band 8C4, was the rarity highlight of our pelagic trip off Half Moon Bay. The bird was banded earlier in 2014 as a nestling in Mexico. Photo by guide Tom Johnson.
PACIFIC LOON (Gavia pacifica) Podicipedidae (Grebes)
PIED-BILLED GREBE (Podilymbus podiceps)
EARED GREBE (Podiceps nigricollis)
WESTERN GREBE (Aechmophorus occidentalis)
CLARK'S GREBE (Aechmophorus clarkii)
Diomedeidae (Albatrosses)
LAYSAN ALBATROSS (Phoebastria immutabilis)
BLACK-FOOTED ALBATROSS (Phoebastria nigripes)
Procellariidae (Shearwaters and Petrels)
NORTHERN FULMAR (Fulmarus glacialis)
PINK-FOOTED SHEARWATER (Puffinus creatopus)

Hawaiian-breeding Black-footed Albatrosses spend a significant portion of their year foraging close to North America in the California Current. We ended up having decent views of these birds, though sometimes they were too close to fit in the camera frame. Photo by guide Tom Johnson.
FLESH-FOOTED SHEARWATER (Puffinus carneipes) SOOTY SHEARWATER (Puffinus griseus)
BLACK-VENTED SHEARWATER (Puffinus opisthomelas)
Hydrobatidae (Storm-Petrels)
WILSON'S STORM-PETREL (Oceanites oceanicus)
ASHY STORM-PETREL (Oceanodroma homochroa)

This Flesh-footed Shearwater was a great addition to a day full of seabirds off Half Moon Bay. It looks mostly like a dark chocolate version of the more common Pink-footed Shearwater. Photo by guide Tom Johnson.
BLACK STORM-PETREL (Oceanodroma melania) Phalacrocoracidae (Cormorants and Shags)
BRANDT'S CORMORANT (Phalacrocorax penicillatus)
DOUBLE-CRESTED CORMORANT (Phalacrocorax auritus)
PELAGIC CORMORANT (Phalacrocorax pelagicus)
Pelecanidae (Pelicans)
AMERICAN WHITE PELICAN (Pelecanus erythrorhynchos)
BROWN PELICAN (Pelecanus occidentalis)
Ardeidae (Herons, Egrets, and Bitterns)

This juvenile Red-shouldered Hawk showed off for us along the East Slope of the Sierra Nevada. Photo by guide Tom Johnson.
GREAT BLUE HERON (Ardea herodias) GREAT EGRET (Ardea alba)
SNOWY EGRET (Egretta thula)
BLACK-CROWNED NIGHT-HERON (Nycticorax nycticorax)
Threskiornithidae (Ibises and Spoonbills)
WHITE-FACED IBIS (Plegadis chihi)
Cathartidae (New World Vultures)
TURKEY VULTURE (Cathartes aura)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus)
Accipitridae (Hawks, Eagles, and Kites)
WHITE-TAILED KITE (Elanus leucurus)
GOLDEN EAGLE (Aquila chrysaetos)
NORTHERN HARRIER (Circus cyaneus)
COOPER'S HAWK (Accipiter cooperii)
BALD EAGLE (Haliaeetus leucocephalus)
The newly split Ridgway's Rail is the western representative of the Clapper Rail complex. We had some excellent studies of these handsome mud hens near the San Francisco airport. Video by guide Chris Benesh.
RED-SHOULDERED HAWK (CALIFORNIA) (Buteo lineatus elegans) RED-TAILED HAWK (Buteo jamaicensis)
Rallidae (Rails, Gallinules, and Coots)
RIDGWAY'S RAIL (Rallus obsoletus)
COMMON GALLINULE (Gallinula galeata)
AMERICAN COOT (Fulica americana)
Recurvirostridae (Stilts and Avocets)
BLACK-NECKED STILT (Himantopus mexicanus)
AMERICAN AVOCET (Recurvirostra americana)
Haematopodidae (Oystercatchers)
BLACK OYSTERCATCHER (Haematopus bachmani)
Charadriidae (Plovers and Lapwings)
BLACK-BELLIED PLOVER (Pluvialis squatarola)
SNOWY PLOVER (Charadrius nivosus)
SEMIPALMATED PLOVER (Charadrius semipalmatus)
KILLDEER (Charadrius vociferus)
Scolopacidae (Sandpipers and Allies)

Black Turnstones were the most widespread "rocky shorebird" of this tour - we saw them at several stops along the coast and in San Francisco Bay. Photo by guide Tom Johnson.
SPOTTED SANDPIPER (Actitis macularius) WANDERING TATTLER (Tringa incana)
GREATER YELLOWLEGS (Tringa melanoleuca)
WILLET (WESTERN) (Tringa semipalmata inornata)
LESSER YELLOWLEGS (Tringa flavipes)
WHIMBREL (AMERICAN) (Numenius phaeopus hudsonicus)
LONG-BILLED CURLEW (Numenius americanus)
MARBLED GODWIT (Limosa fedoa)
RUDDY TURNSTONE (Arenaria interpres)
BLACK TURNSTONE (Arenaria melanocephala)
SANDERLING (Calidris alba)
LEAST SANDPIPER (Calidris minutilla)
PECTORAL SANDPIPER (Calidris melanotos)

A pair of delightful adult Tufted Puffins sat on the water and allowed us to drive up alongside on our Half Moon Bay pelagic trip. Photo by guide Tom Johnson.
WESTERN SANDPIPER (Calidris mauri) SHORT-BILLED DOWITCHER (Limnodromus griseus)
LONG-BILLED DOWITCHER (Limnodromus scolopaceus)
WILSON'S PHALAROPE (Phalaropus tricolor)
RED-NECKED PHALAROPE (Phalaropus lobatus)
Stercorariidae (Skuas and Jaegers)
SOUTH POLAR SKUA (Stercorarius maccormicki)
POMARINE JAEGER (Stercorarius pomarinus)
PARASITIC JAEGER (Stercorarius parasiticus)
LONG-TAILED JAEGER (Stercorarius longicaudus)
Alcidae (Auks, Murres, and Puffins)
COMMON MURRE (Uria aalge)
PIGEON GUILLEMOT (Cepphus columba)
MARBLED MURRELET (Brachyramphus marmoratus)

Elegant Terns, Heermann's Gulls, and Brown Pelicans are all Mexican breeders that move north in the summer, forming a large percentage of the coastal birdlife of central California at this time of year. Photo by guide Tom Johnson.
CASSIN'S AUKLET (Ptychoramphus aleuticus) RHINOCEROS AUKLET (Cerorhinca monocerata)
TUFTED PUFFIN (Fratercula cirrhata)
Laridae (Gulls, Terns, and Skimmers)
SABINE'S GULL (Xema sabini)
HEERMANN'S GULL (Larus heermanni)
RING-BILLED GULL (Larus delawarensis)
WESTERN GULL (Larus occidentalis)
CALIFORNIA GULL (Larus californicus)
GLAUCOUS-WINGED GULL (Larus glaucescens)

This juvenile Long-tailed Jaeger inspired a healthy ID debate during our Half Moon Bay pelagic trip; despite the large amount of white in the upperwing, we confirmed the ID as Long-tailed by the overall shape, light flight style, tiny bill size, rounded central tail points, and the presence of only two white outer primary shafts. Photo by guide Tom Johnson.
CASPIAN TERN (Hydroprogne caspia) COMMON TERN (Sterna hirundo)
FORSTER'S TERN (Sterna forsteri)
ELEGANT TERN (Thalasseus elegans)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
BAND-TAILED PIGEON (Patagioenas fasciata)
EURASIAN COLLARED-DOVE (Streptopelia decaocto) [I]
MOURNING DOVE (Zenaida macroura)
Cuculidae (Cuckoos)
GREATER ROADRUNNER (Geococcyx californianus)
Strigidae (Owls)
BURROWING OWL (Athene cunicularia)
Apodidae (Swifts)
WHITE-THROATED SWIFT (Aeronautes saxatalis)
Trochilidae (Hummingbirds)
ANNA'S HUMMINGBIRD (Calypte anna)
Picidae (Woodpeckers)
LEWIS'S WOODPECKER (Melanerpes lewis)
ACORN WOODPECKER (Melanerpes formicivorus)
WILLIAMSON'S SAPSUCKER (Sphyrapicus thyroideus)

In a burned forest south of Lee Vining, this male Williamson's Sapsucker vied with Least Chipmunks, a Hairy Woodpecker, and a Black-backed Woodpecker for rights to this tree. Photo by guide Tom Johnson.
RED-BREASTED SAPSUCKER (Sphyrapicus ruber) NUTTALL'S WOODPECKER (Picoides nuttallii)
DOWNY WOODPECKER (Picoides pubescens)
HAIRY WOODPECKER (Picoides villosus)
WHITE-HEADED WOODPECKER (Picoides albolarvatus)
BLACK-BACKED WOODPECKER (Picoides arcticus)
NORTHERN FLICKER (Colaptes auratus)
PILEATED WOODPECKER (Dryocopus pileatus)
Falconidae (Falcons and Caracaras)
AMERICAN KESTREL (Falco sparverius)
PRAIRIE FALCON (Falco mexicanus)
Tyrannidae (Tyrant Flycatchers)
OLIVE-SIDED FLYCATCHER (Contopus cooperi)
PACIFIC-SLOPE FLYCATCHER (Empidonax difficilis)

The huge flock of Pinyon Jays in the Jeffrey Pines at Mono Crater was a stunning sight. These hypersocial birds streamed noisily past us over white volcanic soil, and all was wonderful in the world. Photo by guide Tom Johnson.
BLACK PHOEBE (Sayornis nigricans) SAY'S PHOEBE (Sayornis saya)
Laniidae (Shrikes)
LOGGERHEAD SHRIKE (Lanius ludovicianus)
Vireonidae (Vireos)
CASSIN'S VIREO (Vireo cassinii)
HUTTON'S VIREO (Vireo huttoni)
WARBLING VIREO (Vireo gilvus)
Corvidae (Crows, Jays, and Magpies)
PINYON JAY (Gymnorhinus cyanocephalus)
STELLER'S JAY (Cyanocitta stelleri)
WESTERN SCRUB-JAY (COASTAL) (Aphelocoma californica californica)
BLACK-BILLED MAGPIE (Pica hudsonia)
YELLOW-BILLED MAGPIE (Pica nuttalli) [E]
CLARK'S NUTCRACKER (Nucifraga columbiana)
AMERICAN CROW (Corvus brachyrhynchos)
COMMON RAVEN (Corvus corax)
Alaudidae (Larks)
HORNED LARK (Eremophila alpestris)
Hirundinidae (Swallows)
NORTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx serripennis)
TREE SWALLOW (Tachycineta bicolor)
VIOLET-GREEN SWALLOW (Tachycineta thalassina)

At high elevation in the Sierra Nevada and also near Mono Crater, we had some great views of this spectacularly interesting Corvid, the Clark's Nutcracker. Photo by guide Tom Johnson.
BARN SWALLOW (Hirundo rustica) CLIFF SWALLOW (Petrochelidon pyrrhonota)
Paridae (Chickadees and Tits)
MOUNTAIN CHICKADEE (Poecile gambeli)
CHESTNUT-BACKED CHICKADEE (Poecile rufescens)
OAK TITMOUSE (Baeolophus inornatus)
Aegithalidae (Long-tailed Tits)
BUSHTIT (Psaltriparus minimus)
Sittidae (Nuthatches)
RED-BREASTED NUTHATCH (Sitta canadensis)

Several of the species on this tour helped us visualize some quirks of biogeography. We got to see two very different groups of Chestnut-backed Chickadees, including the chestnut-flanked birds of the western Sierra Nevada, part of the northern population, as well as the gray-flanked coastal birds near San Francisco (as shown here). Photo by guide Tom Johnson.
WHITE-BREASTED NUTHATCH (Sitta carolinensis) PYGMY NUTHATCH (Sitta pygmaea)
Certhiidae (Treecreepers)
BROWN CREEPER (Certhia americana)
Troglodytidae (Wrens)
ROCK WREN (Salpinctes obsoletus)
CANYON WREN (Catherpes mexicanus)
HOUSE WREN (Troglodytes aedon)
PACIFIC WREN (SOUTHERN) (Troglodytes pacificus pacificus)
MARSH WREN (PACIFIC) (Cistothorus palustris paludicola)
BEWICK'S WREN (Thryomanes bewickii)
Cinclidae (Dippers)
AMERICAN DIPPER (Cinclus mexicanus)
Regulidae (Kinglets)
GOLDEN-CROWNED KINGLET (Regulus satrapa)
RUBY-CROWNED KINGLET (Regulus calendula)
Paradoxornithidae (Parrotbills, Wrentit, and Allies)
WRENTIT (Chamaea fasciata)
Turdidae (Thrushes and Allies)
WESTERN BLUEBIRD (Sialia mexicana)
MOUNTAIN BLUEBIRD (Sialia currucoides)
TOWNSEND'S SOLITAIRE (Myadestes townsendi)
AMERICAN ROBIN (Turdus migratorius)
Mimidae (Mockingbirds and Thrashers)
CALIFORNIA THRASHER (Toxostoma redivivum)

Under the lush canopy of huge sequoia trees at Calaveras Big Tree State Park, this Pacific Wren belted out its complex, rapid-fire song from atop a dead log. Photo by guide Tom Johnson.
SAGE THRASHER (Oreoscoptes montanus) NORTHERN MOCKINGBIRD (Mimus polyglottos)
Sturnidae (Starlings)
EUROPEAN STARLING (Sturnus vulgaris) [I]
Ptilogonatidae (Silky-flycatchers)
PHAINOPEPLA (Phainopepla nitens)
Parulidae (New World Warblers)
ORANGE-CROWNED WARBLER (Oreothlypis celata)
NASHVILLE WARBLER (WESTERN) (Oreothlypis ruficapilla ridgwayi)
MACGILLIVRAY'S WARBLER (Geothlypis tolmiei)
COMMON YELLOWTHROAT (Geothlypis trichas)
YELLOW WARBLER (Setophaga petechia)
YELLOW-RUMPED WARBLER (AUDUBON'S) (Setophaga coronata auduboni)

Sage Thrashers ran around like shorebirds along the edge of Mono Lake. Along with sparrows, blackbirds, and gulls, the thrashers were attracted by the abundant alkali flies found along the lakeshore. Photo by guide Tom Johnson.
BLACK-THROATED GRAY WARBLER (Setophaga nigrescens) TOWNSEND'S WARBLER (Setophaga townsendi)
HERMIT WARBLER (Setophaga occidentalis)
WILSON'S WARBLER (Cardellina pusilla)
Emberizidae (Buntings and New World Sparrows)
GREEN-TAILED TOWHEE (Pipilo chlorurus)
SPOTTED TOWHEE (Pipilo maculatus)
CALIFORNIA TOWHEE (Melozone crissalis)
CHIPPING SPARROW (Spizella passerina)
BREWER'S SPARROW (Spizella breweri)
VESPER SPARROW (Pooecetes gramineus)
LARK SPARROW (Chondestes grammacus)
BELL'S SPARROW (BELL'S) (Artemisiospiza belli belli)
SAGEBRUSH SPARROW (Artemisiospiza nevadensis)
SAVANNAH SPARROW (Passerculus sandwichensis)
FOX SPARROW (THICK-BILLED) (Passerella iliaca megarhyncha)

We were fortunate to find both Coastal Bell's Sparrows (in the Diablo Range) and interior Sagebrush Sparrows (shown here at Mono Lake) during the tour. Future field work will surely clarify the borders of distribution between these recently split taxa. Photo by guide Tom Johnson.
SONG SPARROW (Melospiza melodia) WHITE-CROWNED SPARROW (MOUNTAIN) (Zonotrichia leucophrys oriantha)
WHITE-CROWNED SPARROW (NUTTALL'S) (Zonotrichia leucophrys nuttalli)
DARK-EYED JUNCO (Junco hyemalis)
Cardinalidae (Cardinals and Allies)
WESTERN TANAGER (Piranga ludoviciana)
BLACK-HEADED GROSBEAK (Pheucticus melanocephalus)
Icteridae (Troupials and Allies)
RED-WINGED BLACKBIRD (Agelaius phoeniceus)
TRICOLORED BLACKBIRD (Agelaius tricolor)
BREWER'S BLACKBIRD (Euphagus cyanocephalus)
BROWN-HEADED COWBIRD (Molothrus ater)
Fringillidae (Siskins, Crossbills, and Allies)
HOUSE FINCH (Haemorhous mexicanus)
CASSIN'S FINCH (Haemorhous cassinii)
RED CROSSBILL (Loxia curvirostra)
PINE SISKIN (Spinus pinus)
LESSER GOLDFINCH (Spinus psaltria)
LAWRENCE'S GOLDFINCH (Spinus lawrencei)
AMERICAN GOLDFINCH (Spinus tristis)
EVENING GROSBEAK (Coccothraustes vespertinus)
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus) [I]
BRUSH RABBIT (Sylvilagus bachmani)
NUTTALL'S (MOUNTAIN) COTTONTAIL (Sylvilagus nuttalli)
DESERT COTTONTAIL (Sylvilagus audubonii)
BLACK-TAILED JACKRABBIT (Lepus californicus)
LEAST CHIPMUNK (Tamias minimus)
MERRIAM'S CHIPMUNK (Tamias merriami)
LONG-EARED CHIPMUNK (Tamias quadrimaculatus)
LODGEPOLE CHIPMUNK (Tamias speciosus)
YELLOW-BELLIED MARMOT (Marmota flaviventris)
CALIFORNIA GROUND SQUIRREL (Spermophilus beecheyi)
GOLDEN-MANTLED GROUND SQUIRREL (Spermophilus lateralis)
FOX SQUIRREL (Sciurus niger)
WESTERN GRAY SQUIRREL (Sciurus griseus)
CHICKAREE (Tamiasciurus douglasii)
BOTTLENOSE DOLPHIN (Tursiops truncatus)
RISSO'S DOLPHIN (Grampus griseus)

One of nine species of marine mammals encountered on our pelagic trip out of Half Moon Bay, this Northern Fur Seal was probably from the population that breeds on the nearby Farallon Islands. Photo by guide Tom Johnson.
HARBOR PORPOISE (Phocoena phocoena) DALL'S PORPOISE (Phocoenoides dalli)
HUMPBACK WHALE (Megaptera novaeangliae)
COYOTE (Canis latrans)
LONG-TAILED WEASEL (Mustela frenata)
CALIFORNIA SEA LION (Zalophus californianus)
STELLER'S SEA LION (Eumetopias jubatus)
NORTHERN FUR SEAL (Callorhinus ursinus)
HARBOR SEAL (Phoca vitulina)
MULE DEER (Odocoileus hemionus)
Totals for the tour: 207 bird taxa and 26 mammal taxa