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Marvelous
Spatuletail at the Huembo feeders: Certainly one of the
prizes of the trip, it was great to see both sexes and
different ages (at least four birds) so well. (Photo by
guide Richard Webster)
We were lucky at the Lomas de Lachay, where in a quick visit
before our flight north we encountered several specialties
(Thick-billed, Coastal, and Grayish miners, Cactus Canastero),
saw some other fine birds (like Least Seedsnipe), and got a
taste of the fog forest of the coastal slope.
In the north, starting at Batan Grande (Bosque de Pomac) we
quickly saw Peruvian Plantcutter, but had to work for Rufous
Flycatcher, also finding Cinereous Finch and enjoying a visit
from the often distant Tumbes Swallows. A quick march up the
beach to the mouth of the Rio Reque was good for many common
waterbirds, and the thrill of a last-second Peruvian Tern
flying by at close range. Our lodge at Chaparri was, as usual,
infested with the Critically Endangered White-winged Guan, and
we had other new birds such as Tumbes Tyrant, Tumbes Sparrow,
and Sulphur-throated Finch.
Quebrada Limon was our campsite for one night, and a Scrub
Nightjar after dinner was a fun find. We split in wider and
shorter ranging groups the next day, both of which saw the
long-term populations of White-winged Guan along with many
Tumbesian species such as White-headed Brush-Finch,
Gray-and-gold Warbler and Pacific Elaenia, while having
divided fortunes on some others (Ecuadorian Trogon and Tumbes
Hummingbird, which was especially scarce this year). Before
leaving the Pacific slope, we birded for a morning on Abra
Porculla, finding Piura Chat-Tyrant, and, with a morning of
looking inside bushes, Henna-hooded and Rufous-necked
Foliage-gleaners, Bay-crowned Brush-Finch, Black-cowled
Saltator, and Ecuadorian Piculet.
Based in Jaen we started our survey of the arid habitats of
the Maranon, starting with the lovely Maranon Crescentchest
and continuing with such skulkers as Chichipe Spinetail
(fairly easy) and Maranon Spinetail (awful), leaving the
following morning with improved views of Little Inca-Finch
before heading into the Eastern Andes.
Climbing above the Rio Utcubamba (Fasciated Tiger-Heron), we
reached the Huembo Spatuletail Center, where Santos' feeders
were fortunately hosting Spatuletails, including two adult
males, along with a nice variety of other hummingbirds. The
next morning we split up along the Rio Chido, with an
energetic group making the hike well up the valley to find a
great bird, the Pale-billed Antpitta, while the other group
had a really sloooow morning in the same forest that had been
busy the evening before (White-capped Tanager was nice).
Leaving those paintings behind for four nights, we went to
the wilds of Abra Patricia and Owlet Lodge. Two attempts for
the owlet produced muddy boots (at the least!) and an owlet
that was heard and heard, but not seen. However, our days
there and down the mountain to the edge of Amazonia were rich
indeed, and highlights were many, ranging from Yellow-scarfed
Tanagers near the lodge, the surprise of a cooperative
Ash-throated Antwren, good views of Royal Sunangel and
Bar-winged Wood-Wren (after several attempts) in the stunted
forest, several antpittas (but fewer seen by fewer people than
normal), the Little Darling Gem of a Tody-Monster, more
dramatic specialties like Black-bellied and Yellow-crested
tanagers and less dramatic endemics like Inca Flycatcher and
Speckle-chested Piculet, and a pile of foothills specialties
(Equatorial Graytail, Vermilion, Blue-browed, and Golden-eared
tanagers, Ecuadorian Tyrannulet, Golden-collared Honeycreeper)
and just plain great birds (Andean Cock-of-the-Rock for
starters, and those Lanceolated Monklets!). And the moths at
the Lodge! And the Emerald-bellied Pufflegs at the feeders!
And best of all, that wonderful troop of Yellow-tailed Woolly
Monkeys, perhaps the hardest to replicate find of the trip.
Heading back up the Utcubamba Valley, we saw our first
Peruvian Pigeons and Black-necked Woodpeckers, with a bonus
flock of Andean Parakeets. We were again lucky with the day
roost of Koepcke's Screech-Owls. The Leymebamba museum was a
treat, and the nearby feeders busy (Sword-billed, Rainbow
Starfrontlet, Purple-throated Sunangel), while the forest up
canyon had Gray-breasted Mountain-Toucan and Golden-headed
Quetzal. Two mornings on Abra Barro Negro were productive,
including Russet-mantled Softtail, our first species of
"Rufous" Antpitta, Coppery Metaltail, and Golden-plumed
Parakeets among many highland species.
We spent hours searching for Yellow-faced Parrotlet on both
sides of our camp at Balsas, finding two single birds, but
ending with exciting views for everyone of this missable bird.
In the process we also had excellent studies of Buff-bridled
Inca-Finch after finding our first Chestnut-backed Thornbirds.
Moving upslope to Hacienda Limon, we saw our third inca-finch,
the Gray-winged, and then continued to higher elevations and
every more dramatic views. Patches of scrub were home to
Jelski's Chat-Tyrant and Black-crested Tit-Tyrant as well as a
bonus of Plain-tailed Warbling-Finch. Celendin had hot water
showers after our night of camping, and provided a base to
journey onward toward Cajamarca the next morning, finding our
second type of "Rufous" Antpitta, both White-tailed and
Black-billed shrike-tyrants, and the distinctive Striated
Earthcreeper.
We ended our journey with two forays out from Cajamarca, the
first toward San Marcos, where we saw the distinctive Great
Spinetail very well (whew!) and then along the Rio Chonta,
where we found a Gray-bellied Comet (whew!). With some extra
time before our flight, we added our last endemic, the less
than dramatic (but very local) Unicolored Tapaculo.
In addition to being a spectacular avifauna, it is a
threatened one. As a rough count, we encountered 2 Critically
Endangered, 12 Endangered, 21 Vulnerable, and 25 Near
Threatened (seven heard-only overall) based on the
publications of BirdLife International. That is our highest
such total, not because we did so much better, but because
BirdLife keeps adding more to the lists, believing the news
from the front is not so good. We know you give to many
charities, but if your trip here has developed a special
interest, contributions to ECOAN's Abra Patricia and other
reserves can be channeled through the American Bird
Conservancy.
Our journey would not have been possible without the superb
crew from Manu, special thanks to their chief and our chef,
Aurelio, along with Rene, Walter, Alejandro, and Ruben.
Taxonomy follows Clements in large part. We have indicated
other taxa of interest, with additional comments from Handbook
of the Birds of the World (HBW) and the South American
Classification Committee (S.A.C.C.), the excellent new field
guide to Peru, and various journal articles. Apologies are
offered to the Spanish language because we avoid the
orthographic marks that do not translate well across various
computer platforms and programs.
--Richard
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for
individual species as appropriate: * = heard only, I =
introduced, E = endemic, N = nesting, a = austral migrant, b =
boreal migrant
Tinamidae (Tinamous)

A Torrent Duck
family of four was along the Rio Aguas Verdes. (Photo by
guide Richard Webster)
HIGHLAND TINAMOU
(Nothocercus bonapartei) [*]
BROWN TINAMOU (Crypturellus
obsoletus castaneus) [*]
PALE-BROWED TINAMOU (Crypturellus transfasciatus) [*]
TATAUPA TINAMOU (Crypturellus tataupa inops) [*]
ANDEAN TINAMOU (Nothoprocta
pentlandii ambigua)
Anatidae (Ducks, Geese, and Waterfowl)
COMB DUCK (Sarkidiornis
melanotos)
TORRENT DUCK (Merganetta
armata)
CINNAMON TEAL (Anas
cyanoptera)
WHITE-CHEEKED PINTAIL (Anas bahamensis)
RUDDY DUCK (ANDEAN) (Oxyura jamaicensis ferruginea)
Cracidae (Guans, Chachalacas, and Curassows)
SPECKLED CHACHALACA (Ortalis guttata)
ANDEAN GUAN (Penelope
montagnii)
WHITE-WINGED GUAN (Penelope albipennis)
[E]
WATTLED GUAN (Aburria
aburri) [*]
SICKLE-WINGED GUAN (Chamaepetes goudotii)
Odontophoridae (New World Quail)
RUFOUS-BREASTED WOOD-QUAIL (Odontophorus speciosus) [*]
Podicipedidae (Grebes)
WHITE-TUFTED GREBE (Rollandia rolland chilensis)
PIED-BILLED GREBE (Podilymbus podiceps)

Gathering at dawn
to hike up Quebrada Limon, where we had our second
encounter with White-winged Guan. (Photo by guide Richard
Webster)
GREAT GREBE
(Podiceps major)
Sulidae (Boobies and Gannets)
PERUVIAN BOOBY (Sula
variegata)
Phalacrocoracidae (Cormorants and Shags)
NEOTROPIC CORMORANT (Phalacrocorax brasilianus)
Pelecanidae (Pelicans)
PERUVIAN PELICAN (Pelecanus thagus)
Ardeidae (Herons, Egrets, and Bitterns)
FASCIATED TIGER-HERON (Tigrisoma fasciatum)
COCOI HERON (Ardea
cocoi)
GREAT EGRET (Ardea
alba)
SNOWY EGRET (Egretta
thula)
LITTLE BLUE HERON (Egretta caerulea)
CATTLE EGRET (Bubulcus
ibis)
STRIATED HERON (Butorides
striata)

Peruvian Pelicans
on the beach near Puerto Eten, one of the several Humboldt
Current species we saw here. (Photo by guide Richard
Webster)
BLACK-CROWNED
NIGHT-HERON (Nycticorax
nycticorax)
Threskiornithidae (Ibises and Spoonbills)
PUNA IBIS (Plegadis
ridgwayi)
Cathartidae (New World Vultures)
BLACK VULTURE (Coragyps
atratus)
TURKEY VULTURE (Cathartes
aura)
GREATER YELLOW-HEADED VULTURE (Cathartes melambrotus)
KING VULTURE (Sarcoramphus
papa)
Pandionidae (Osprey)
OSPREY (Pandion
haliaetus) [b]
Accipitridae (Hawks, Eagles, and Kites)
HOOK-BILLED KITE (Chondrohierax uncinatus)
PEARL KITE (Gampsonyx
swainsonii)
BLACK-CHESTED BUZZARD-EAGLE (Geranoaetus melanoleucus)
SAVANNA HAWK (Buteogallus
meridionalis)
HARRIS'S HAWK (Parabuteo
unicinctus)
ROADSIDE HAWK (Buteo
magnirostris)

This is as close
as we got to Andean Condor, these three figures in
Codormarca; we have seen Andean Condor on this trip, but
not often, King Vulture being a better prospect. (Photo by
guide Richard Webster)
BROAD-WINGED HAWK
(Buteo platypterus) [b]
VARIABLE HAWK (Buteo
polyosoma)
Falconidae (Falcons and Caracaras)
MOUNTAIN CARACARA (Phalcoboenus megalopterus)
CRESTED CARACARA (Caracara cheriway)
LAUGHING FALCON (Herpetotheres cachinnans)
AMERICAN KESTREL (Falco sparverius)
PEREGRINE FALCON (Falco peregrinus)
Rallidae (Rails, Gallinules, and Coots)
BLACKISH RAIL (Pardirallus
nigricans)
PLUMBEOUS RAIL (Pardirallus
sanguinolentus)
PURPLE GALLINULE (Porphyrio martinica)
COMMON GALLINULE (Gallinula galeata)
SLATE-COLORED COOT (Fulica ardesiaca)
Burhinidae (Thick-knees)
PERUVIAN THICK-KNEE (Burhinus superciliaris)
Charadriidae (Plovers and Lapwings)
ANDEAN LAPWING (Vanellus
resplendens)
BLACK-BELLIED PLOVER (Pluvialis squatarola) [b]
SNOWY PLOVER (PERUVIAN) (Charadrius nivosus occidentalis)
SEMIPALMATED PLOVER (Charadrius semipalmatus) [b]
KILLDEER (Charadrius
vociferus)
Haematopodidae (Oystercatchers)
AMERICAN OYSTERCATCHER (Haematopus palliatus)
Recurvirostridae (Stilts and Avocets)
BLACK-NECKED STILT (BLACK-NECKED) (Himantopus mexicanus mexicanus)
Scolopacidae (Sandpipers and Allies)
SPOTTED SANDPIPER (Actitis macularius)
[b]
GREATER YELLOWLEGS (Tringa melanoleuca) [b]
WILLET (Tringa
semipalmata) [b]
LESSER YELLOWLEGS (Tringa flavipes) [b]
WHIMBREL (Numenius
phaeopus) [b]
HUDSONIAN GODWIT (Limosa haemastica) [b]
RUDDY TURNSTONE (Arenaria interpres) [b]

Peruvian Tern is
a highly endangered species, and one that we generally
miss, so this bird south of Chiclayo was a thrill. (Photo
by guide Richard Webster)
SANDERLING
(Calidris alba) [b]
SEMIPALMATED SANDPIPER (Calidris pusilla)
[b]
LEAST SANDPIPER (Calidris minutilla) [b]
WILSON'S PHALAROPE (Phalaropus tricolor) [b]
Thinocoridae (Seedsnipes)
LEAST SEEDSNIPE (Thinocorus rumicivorus)
Laridae (Gulls, Terns, and Skimmers)
ANDEAN GULL (Chroicocephalus
serranus)
GRAY-HOODED GULL (Chroicocephalus cirrocephalus)
FRANKLIN'S GULL (Leucophaeus pipixcan) [b]
BELCHER'S GULL (Larus
belcheri)
KELP GULL (Larus
dominicanus)
PERUVIAN TERN (Sternula
lorata)
GULL-BILLED TERN (Gelochelidon nilotica) [b]
ELEGANT TERN (Thalasseus
elegans)
[b]
Columbidae (Pigeons and Doves)

Bare-faced
Ground-Dove is one of the short list of species that we
found during our parrotlet searches above the Rio Maranon;
our camp near Balsas was along the river behind the hill
in the center. (Photo by guide Richard Webster)
ROCK PIGEON
(Columba livia) [I]
BAND-TAILED PIGEON (Patagioenas fasciata)
PERUVIAN PIGEON (Patagioenas oenops)
PLUMBEOUS PIGEON (Patagioenas plumbea) [*]
RUDDY PIGEON (Patagioenas
subvinacea)
[*]
WEST PERUVIAN DOVE (Zenaida meloda)
EARED DOVE (Zenaida
auriculata)
ECUADORIAN GROUND-DOVE (Columbina buckleyi)
CROAKING GROUND-DOVE (Columbina cruziana)
BLUE GROUND-DOVE (Claravis pretiosa)
BARE-FACED GROUND-DOVE (Metriopelia ceciliae)
WHITE-TIPPED DOVE (Leptotila verreauxi)
WHITE-THROATED QUAIL-DOVE (Geotrygon frenata)
Psittacidae (Parrots)

This portion of
the Utcubamba Valley was good birding, with a surprise
flock of Andean Parakeets, our first Black-necked
Woodpecker, and another Maranon Thrush. (Photo by guide
Richard Webster)
GOLDEN-PLUMED
PARAKEET (Leptosittaca
branickii)
SCARLET-FRONTED PARAKEET (Aratinga wagleri)
MITRED PARAKEET (Aratinga mitrata)
RED-MASKED PARAKEET (Aratinga erythrogenys)
WHITE-EYED PARAKEET (Aratinga leucophthalma)
ANDEAN PARAKEET (Bolborhynchus orbygnesius)
PACIFIC PARROTLET (Forpus coelestis)

Yellow-faced
Parrotlet was a struggle (as usual), with a happy ending
for all. (Photo by guide Richard Webster)
YELLOW-FACED
PARROTLET (Forpus xanthops)
[E]
BLUE-HEADED PARROT (Pionus menstruus)
RED-BILLED PARROT (Pionus sordidus)
SPECKLE-FACED PARROT (WHITE-CAPPED) (Pionus tumultuosus seniloides)
SCALY-NAPED PARROT (Amazona mercenaria)
Cuculidae (Cuckoos)
SQUIRREL CUCKOO (Piaya cayana)
STRIPED CUCKOO (Tapera
naevia)
SMOOTH-BILLED ANI (Crotophaga ani)
GROOVE-BILLED ANI (Crotophaga sulcirostris)
Strigidae (Owls)
PERUVIAN SCREECH-OWL (Megascops roboratus)
KOEPCKE'S SCREECH-OWL (Megascops koepckeae) [E]
ANDEAN PYGMY-OWL (Glaucidium jardinii)
[*]
FERRUGINOUS PYGMY-OWL (Glaucidium brasilianum) [*]
PERUVIAN PYGMY-OWL (Glaucidium peruanum)

Long-whiskered
Owlet lives down there, and down there we went, hearing it
very well, but not getting a view. This is the view from
Owlet Lodge, and a bird-rich view it is. (Photo by guide
Richard Webster)
LONG-WHISKERED
OWLET (Xenoglaux loweryi)
[E*]
BURROWING OWL (Athene
cunicularia)
RUFOUS-BANDED OWL (Ciccaba albitarsis) [*]
Caprimulgidae (Nightjars and Allies)
LESSER NIGHTHAWK (Chordeiles acutipennis)
SCRUB NIGHTJAR (Caprimulgus
anthonyi)
SWALLOW-TAILED NIGHTJAR (Uropsalis segmentata)
Nyctibiidae (Potoos)
COMMON POTOO (Nyctibius
griseus)
[*]
Apodidae (Swifts)
CHESTNUT-COLLARED SWIFT (Streptoprocne rutila)
WHITE-COLLARED SWIFT (Streptoprocne zonaris)
CHIMNEY SWIFT (Chaetura
pelagica) [b]
SHORT-TAILED SWIFT (Chaetura brachyura)
SHORT-TAILED SWIFT (TUMBES) (Chaetura brachyura ocypetes)
GRAY-RUMPED SWIFT (Chaetura cinereiventris)
WHITE-TIPPED SWIFT (Aeronautes montivagus)
ANDEAN SWIFT (Aeronautes
andecolus)
Trochilidae (Hummingbirds)
GREEN HERMIT (Phaethornis
guy)
TAWNY-BELLIED HERMIT (Phaethornis syrmatophorus) [*]
GRAY-CHINNED HERMIT (Phaethornis griseogularis)
GRAY-CHINNED HERMIT (Phaethornis griseogularis porcullae)
GREEN VIOLETEAR (Colibri thalassinus)
SPARKLING VIOLETEAR (Colibri coruscans)
AMETHYST-THROATED SUNANGEL (Heliangelus amethysticollis)

Purple-throated
Sunangel is a gem, as we saw at Leymebamba; but it has
become a complicated gem, with a possible split, and the
magenta gorget and blue spots on the breast are more like
the one that should not be in Leymebamba??? (Photo by
guide Richard Webster)
PURPLE-THROATED
SUNANGEL (Heliangelus viola)
ROYAL SUNANGEL (Heliangelus
regalis)
WIRE-CRESTED THORNTAIL (Discosura popelairii)
SPECKLED HUMMINGBIRD (Adelomyia melanogenys)
LONG-TAILED SYLPH (Aglaiocercus kingi)
GRAY-BELLIED COMET (Taphrolesbia griseiventris) [E]
BLACK-TAILED TRAINBEARER (Lesbia victoriae)
GREEN-TAILED TRAINBEARER (Lesbia nuna)
RUFOUS-CAPPED THORNBILL (Chalcostigma ruficeps)
TYRIAN METALTAIL (Metallura tyrianthina)
COPPERY METALTAIL (Metallura theresiae) [E]
BLACK METALTAIL (Metallura phoebe) [E]

Marvelous
Spatuletail is larger than a bee, but not all that much
larger! The hummingbirds were bothered by the abundant
bees, but still managed to spend much time at the feeders.
(Photo by guide Richard Webster)
SAPPHIRE-VENTED
PUFFLEG (COPPERY-NAPED) (Eriocnemis
luciani catharina)
EMERALD-BELLIED PUFFLEG (Eriocnemis alinae)
MARVELOUS SPATULETAIL (Loddigesia mirabilis) [E]
SHINING SUNBEAM (Aglaeactis cupripennis)
BRONZY INCA (Coeligena
coeligena)
COLLARED INCA (Coeligena
torquata)
VIOLET-THROATED STARFRONTLET (Coeligena violifer)
RAINBOW STARFRONTLET (Coeligena iris)
MOUNTAIN VELVETBREAST (Lafresnaya lafresnayi)
SWORD-BILLED HUMMINGBIRD (Ensifera ensifera)
GREAT SAPPHIREWING (Pterophanes cyanopterus)
CHESTNUT-BREASTED CORONET (Boissonneaua matthewsii)
FAWN-BREASTED BRILLIANT (Heliodoxa rubinoides)
VIOLET-FRONTED BRILLIANT (Heliodoxa leadbeateri)

Rainbow
Starfrontlet does have a rainbow, at the right angle, an
angle that was available at Adriana's feeders above
Leymebamba. (Photo by guide Richard Webster)
GIANT HUMMINGBIRD
(Patagona gigas)
LONG-BILLED STARTHROAT (Heliomaster longirostris)
PURPLE-COLLARED WOODSTAR (Myrtis fanny)
OASIS HUMMINGBIRD (Rhodopis vesper)
PERUVIAN SHEARTAIL (Thaumastura cora)
WHITE-BELLIED WOODSTAR (Chaetocercus mulsant)
LITTLE WOODSTAR (Chaetocercus bombus)
SHORT-TAILED WOODSTAR (Myrmia micrura)
FORK-TAILED WOODNYMPH (Thalurania furcata)
MANY-SPOTTED HUMMINGBIRD (Taphrospilus hypostictus)
TUMBES HUMMINGBIRD (Leucippus baeri)
SPOT-THROATED HUMMINGBIRD (Leucippus taczanowskii)
[E]
WHITE-BELLIED HUMMINGBIRD (Amazilia chionogaster)
AMAZILIA HUMMINGBIRD (Amazilia amazilia)
ANDEAN EMERALD (Amazilia
franciae)
Trogonidae (Trogons)
GOLDEN-HEADED QUETZAL (Pharomachrus auriceps)
ECUADORIAN TROGON (Trogon mesurus)
Momotidae (Motmots)
WHOOPING MOTMOT (ARGENTICINCTUS) (Momotus subrufescens argenticinctus)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)
AMAZON KINGFISHER (Chloroceryle amazona)
Bucconidae (Puffbirds)
LANCEOLATED MONKLET (Micromonacha lanceolata)
WHITE-FACED NUNBIRD (Hapaloptila castanea)
Capitonidae (New World Barbets)
GILDED BARBET (Capito
auratus) [*]
VERSICOLORED BARBET (Eubucco versicolor)
Ramphastidae (Toucans)
EMERALD TOUCANET (BLACK-THROATED) (Aulacorhynchus prasinus cyanolaemus)
CHESTNUT-TIPPED TOUCANET (Aulacorhynchus derbianus)
GRAY-BREASTED MOUNTAIN-TOUCAN (Andigena hypoglauca)
BLACK-MANDIBLED TOUCAN (Ramphastos ambiguus) [*]
Picidae (Woodpeckers)
ECUADORIAN PICULET (Picumnus sclateri)
SPECKLE-CHESTED PICULET (Picumnus steindachneri) [E]
YELLOW-TUFTED WOODPECKER (Melanerpes cruentatus)
SMOKY-BROWN WOODPECKER (Picoides fumigatus)
LITTLE WOODPECKER (Veniliornis passerinus)
SCARLET-BACKED WOODPECKER (Veniliornis callonotus)
GOLDEN-OLIVE WOODPECKER (Colaptes rubiginosus)
BLACK-NECKED WOODPECKER (Colaptes atricollis) [E]
ANDEAN FLICKER (Colaptes
rupicola)
LINEATED WOODPECKER (Dryocopus lineatus)
GUAYAQUIL WOODPECKER (Campephilus gayaquilensis) [*]
Furnariidae (Ovenbirds and Woodcreepers)
COASTAL MINER (Geositta
peruviana) [E]
GRAYISH MINER (Geositta
maritima)
THICK-BILLED MINER (Geositta crassirostris) [E]
STRIATED EARTHCREEPER (Geocerthia serrana) [E]
CREAM-WINGED CINCLODES (Cinclodes albiventris)
WHITE-WINGED CINCLODES (Cinclodes atacamensis)
PALE-LEGGED HORNERO (PACIFIC) (Furnarius leucopus cinnamomeus)
WREN-LIKE RUSHBIRD (Phleocryptes melanops)
RUSTY-CROWNED TIT-SPINETAIL (Leptasthenura pileata cajabambae) [E]
AZARA'S SPINETAIL (Synallaxis azarae)
DARK-BREASTED SPINETAIL (Synallaxis albigularis)
RUFOUS SPINETAIL (Synallaxis unirufa)
MARANON SPINETAIL (Synallaxis maranonica)
NECKLACED SPINETAIL (NECKLACED) (Synallaxis stictothorax maculata)
NECKLACED SPINETAIL (CHINCHIPE) (Synallaxis stictothorax chinchipensis)

Great Spinetail
(Photo by guide Richard Webster)
GREAT SPINETAIL
(Siptornopsis hypochondriaca) [E]
ASH-BROWED SPINETAIL (Cranioleuca curtata)
LINE-CHEEKED SPINETAIL (Cranioleuca antisiensis)
BARON'S SPINETAIL (Cranioleuca baroni)
[E]
RUSSET-MANTLED SOFTTAIL (Thripophaga berlepschi) [E]
MANY-STRIPED CANASTERO (Asthenes flammulata)
STREAK-THROATED CANASTERO (Asthenes humilis)
WHITE-CHINNED THISTLETAIL (Asthenes fuliginosa peruviana)
CACTUS CANASTERO (Pseudasthenes cactorum) [E]
RUFOUS-FRONTED THORNBIRD (RUFOUS-FRONTED)
(Phacellodomus rufifrons peruvianus)
CHESTNUT-BACKED THORNBIRD (Phacellodomus dorsalis) [E]
EQUATORIAL GRAYTAIL (Xenerpestes singularis)
PEARLED TREERUNNER (Margarornis squamiger)
STREAKED TUFTEDCHEEK (Pseudocolaptes boissonneautii)
MONTANE FOLIAGE-GLEANER (Anabacerthia striaticollis)
RUFOUS-NECKED FOLIAGE-GLEANER (Syndactyla ruficollis)
BUFF-FRONTED FOLIAGE-GLEANER (Philydor rufum)
STRIPED TREEHUNTER (Thripadectes holostictus)
HENNA-HOODED FOLIAGE-GLEANER (Hylocryptus erythrocephalus)
STREAKED XENOPS (Xenops rutilans)
TYRANNINE WOODCREEPER (Dendrocincla tyrannina)
OLIVACEOUS WOODCREEPER (Sittasomus griseicapillus)
STRONG-BILLED WOODCREEPER (Xiphocolaptes promeropirhynchus)
OLIVE-BACKED WOODCREEPER (Xiphorhynchus triangularis)
STREAK-HEADED WOODCREEPER (Lepidocolaptes souleyetii)
MONTANE WOODCREEPER (Lepidocolaptes lacrymiger)
Thamnophilidae (Typical Antbirds)
RUFOUS-CAPPED ANTSHRIKE (Thamnophilus ruficapillus jaczewskii)
CHAPMAN'S ANTSHRIKE (Thamnophilus zarumae)
LINED ANTSHRIKE (Thamnophilus tenuepunctatus)
COLLARED ANTSHRIKE (Thamnophilus bernardi)
COLLARED ANTSHRIKE (SHUMBAE) (Thamnophilus bernardi shumbae) [*]
NORTHERN SLATY-ANTSHRIKE (MARANON) (Thamnophilus punctatus leucogaster)
VARIABLE ANTSHRIKE (Thamnophilus caerulescens subandinus)
RUSSET ANTSHRIKE (Thamnistes anabatinus)
PLAIN ANTVIREO (Dysithamnus
mentalis)
SLATY ANTWREN (Myrmotherula
schisticolor)
ASH-THROATED ANTWREN (Herpsilochmus parkeri) [E]
YELLOW-BREASTED ANTWREN (Herpsilochmus axillaris)
LONG-TAILED ANTBIRD (Drymophila caudata)
RUFOUS-RUMPED ANTWREN (Terenura callinota)
BLACKISH ANTBIRD (Cercomacra nigrescens)
WHITE-BACKED FIRE-EYE (Pyriglena leuconota)
Formicariidae (Antthrushes)
BARRED ANTTHRUSH (Chamaeza mollissima)
[*]
Grallariidae (Antpittas)
UNDULATED ANTPITTA (Grallaria squamigera) [*]
CHESTNUT-CROWNED ANTPITTA (Grallaria ruficapilla)
PALE-BILLED ANTPITTA (Grallaria carrikeri) [E]
RUSTY-TINGED ANTPITTA (Grallaria przewalskii) [E]

Rufous Antpitta
was one of several specialties we saw on our ascent of
Abra Barro Negro; here the group scans the forest for
Sapphire-vented Puffleg and White-chinned Thisteltail.
(Photo by guide Richard Webster)
RUFOUS ANTPITTA
(Grallaria rufula cajamarcae)
RUFOUS ANTPITTA (Grallaria rufula obscura)
CHESTNUT ANTPITTA (Grallaria blakei) [E]
OCHRE-FRONTED ANTPITTA (Grallaricula ochraceifrons) [E*]
RUSTY-BREASTED ANTPITTA (TUMBES) (Grallaricula ferrugineipectus leymebambae)
[*]
Conopophagidae (Gnateaters)
CHESTNUT-CROWNED GNATEATER (Conopophaga castaneiceps) [*]
Rhinocryptidae (Tapaculos)
ASH-COLORED TAPACULO (Myornis senilis)
BLACKISH TAPACULO (PERUVIAN) (Scytalopus latrans intermedius)
UNICOLORED TAPACULO (Scytalopus unicolor) [E]
TRILLING TAPACULO (Scytalopus parvirostris)
RUFOUS-VENTED TAPACULO (Scytalopus femoralis) [E]
WHITE-CROWNED TAPACULO (Scytalopus atratus atratus)
Melanopareiidae (Crescentchests)
MARANON CRESCENTCHEST (Melanopareia maranonica)
ELEGANT CRESCENTCHEST (Melanopareia elegans)
Tyrannidae (Tyrant Flycatchers)
SOUTHERN BEARDLESS-TYRANNULET (Camptostoma obsoletum)
WHITE-BANDED TYRANNULET (Mecocerculus stictopterus)
WHITE-THROATED TYRANNULET (Mecocerculus leucophrys)
SULPHUR-BELLIED TYRANNULET (Mecocerculus minor)

Black-crested
Tit-Tyrant and Jelski's Chat-Tyrant were ahead of us as we
climbed above Balsas and Hacienda Limon. (Photo by guide
Richard Webster)
BLACK-CRESTED
TIT-TYRANT (Anairetes
nigrocristatus)
YELLOW-BILLED TIT-TYRANT (Anairetes flavirostris)
TUFTED TIT-TYRANT (Anairetes parulus)
MOUSE-COLORED TYRANNULET (TUMBES) (Phaeomyias murina tumbezana)
PACIFIC ELAENIA (Myiopagis subplacens)
YELLOW-BELLIED ELAENIA (Elaenia flavogaster)
WHITE-CRESTED ELAENIA (Elaenia albiceps)
MOTTLE-BACKED ELAENIA (Elaenia gigas)
SIERRAN ELAENIA (Elaenia pallatangae)
TORRENT TYRANNULET (Serpophaga cinerea)
STREAK-NECKED FLYCATCHER (Mionectes striaticollis)
OLIVE-STRIPED FLYCATCHER (Mionectes olivaceus)
SLATY-CAPPED FLYCATCHER (Leptopogon superciliaris)
INCA FLYCATCHER (Leptopogon taczanowskii) [E]
MARBLE-FACED BRISTLE-TYRANT (Phylloscartes ophthalmicus)
MOTTLE-CHEEKED TYRANNULET (Phylloscartes ventralis)
ECUADORIAN TYRANNULET (Phylloscartes gualaquizae)
BLACK-CAPPED TYRANNULET (Phyllomyias nigrocapillus)
ASHY-HEADED TYRANNULET (Phyllomyias cinereiceps) [*]
TAWNY-RUMPED TYRANNULET (Phyllomyias uropygialis)
PLUMBEOUS-CROWNED TYRANNULET (Phyllomyias plumbeiceps)
GOLDEN-FACED TYRANNULET (Zimmerius chrysops)
ORNATE FLYCATCHER (Myiotriccus ornatus)
MANY-COLORED RUSH TYRANT (Tachuris rubrigastra)
RUFOUS-HEADED PYGMY-TYRANT (Pseudotriccus ruficeps)
TAWNY-CROWNED PYGMY-TYRANT (Euscarthmus meloryphus)
GRAY-AND-WHITE TYRANNULET (Pseudelaenia leucospodia)
SCALE-CRESTED PYGMY-TYRANT (Lophotriccus pileatus)
BLACK-THROATED TODY-TYRANT (Hemitriccus granadensis)
JOHNSON'S TODY-FLYCATCHER (Poecilotriccus luluae) [E]
COMMON TODY-FLYCATCHER (Todirostrum cinereum)
YELLOW-OLIVE FLYCATCHER (Tolmomyias sulphurescens)
CINNAMON FLYCATCHER (Pyrrhomyias cinnamomeus)
OLIVE-CHESTED FLYCATCHER (Myiophobus cryptoxanthus)
BRAN-COLORED FLYCATCHER (Myiophobus fasciatus)
GRAY-BREASTED FLYCATCHER (Lathrotriccus griseipectus)
OLIVE-SIDED FLYCATCHER (Contopus cooperi) [b]
SMOKE-COLORED PEWEE (Contopus fumigatus)
WESTERN WOOD-PEWEE (Contopus sordidulus) [b]
TROPICAL PEWEE (TUMBES) (Contopus cinereus punensis)
ALDER FLYCATCHER (Empidonax alnorum) [b]
BLACK PHOEBE (Sayornis
nigricans)
VERMILION FLYCATCHER (Pyrocephalus rubinus)
RUFOUS-TAILED TYRANT (Knipolegus poecilurus)
RUFOUS-NAPED GROUND-TYRANT (Muscisaxicola rufivertex)
BLACK-BILLED SHRIKE-TYRANT (Agriornis montanus)

White-tailed
Shrike-Tyrant is one of the species we saw near
Hillstarless Hill west of Celendin; there were probably
some hillstars somewhere out there, but they were not at
our site, where there were few flowers. (Photo by guide
Richard Webster)
WHITE-TAILED
SHRIKE-TYRANT (Agriornis
albicauda)
STREAK-THROATED BUSH-TYRANT (Myiotheretes striaticollis)
RUFOUS-WEBBED BUSH-TYRANT (Polioxolmis rufipennis)
TUMBES TYRANT (Tumbezia
salvini)
JELSKI'S CHAT-TYRANT (Ochthoeca jelskii)
SLATY-BACKED CHAT-TYRANT (MAROON-BELTED)
(Ochthoeca cinnamomeiventris
angustifasciata)
RUFOUS-BREASTED CHAT-TYRANT (Ochthoeca rufipectoralis)
BROWN-BACKED CHAT-TYRANT (Ochthoeca fumicolor)
PIURA CHAT-TYRANT (Ochthoeca piurae) [E]
WHITE-BROWED CHAT-TYRANT (Ochthoeca leucophrys)
SHORT-TAILED FIELD TYRANT (Muscigralla brevicauda)
RUFOUS FLYCATCHER (Myiarchus semirufus) [E]
DUSKY-CAPPED FLYCATCHER (Myiarchus tuberculifer atriceps)

Evening at camp
in Balsas, and time for the list with beer and wine.
(Photo by guide Richard Webster)
SOOTY-CROWNED
FLYCATCHER (Myiarchus
phaeocephalus phaeocephalus)
SOOTY-CROWNED FLYCATCHER (Myiarchus phaeocephalus interior)
PALE-EDGED FLYCATCHER (Myiarchus cephalotes)
BROWN-CRESTED FLYCATCHER (Myiarchus tyrannulus)
GREAT KISKADEE (Pitangus
sulphuratus)
BOAT-BILLED FLYCATCHER (Megarynchus pitangua) [*]
SOCIAL FLYCATCHER (Myiozetetes similis)
BAIRD'S FLYCATCHER (Myiodynastes bairdii)
PIRATIC FLYCATCHER (Legatus leucophaius)
TROPICAL KINGBIRD (Tyrannus melancholicus)
Cotingidae (Cotingas)
GREEN-AND-BLACK FRUITEATER (Pipreola riefferii)
BARRED FRUITEATER (Pipreola arcuata) [*]
PERUVIAN PLANTCUTTER (Phytotoma raimondii) [E]
RED-CRESTED COTINGA (Ampelion rubrocristatus)
CHESTNUT-CRESTED COTINGA (Ampelion rufaxilla)
ANDEAN COCK-OF-THE-ROCK (Rupicola peruvianus)
Pipridae (Manakins)
WING-BARRED PIPRITES (Piprites chloris)
Tityridae (Tityras and Allies)
MASKED TITYRA (Tityra
semifasciata)
GREEN-BACKED BECARD (YELLOW-CHEEKED)
(Pachyramphus viridis xanthogenys)
BARRED BECARD (Pachyramphus
versicolor)
BLACK-AND-WHITE BECARD (Pachyramphus albogriseus)
Vireonidae (Vireos)
BROWN-CAPPED VIREO (Vireo leucophrys) [*]
RED-EYED VIREO (Vireo
olivaceus)
OLIVACEOUS GREENLET (Hylophilus olivaceus)
RUFOUS-BROWED PEPPERSHRIKE (Cyclarhis gujanensis)
Corvidae (Crows, Jays, and Magpies)
WHITE-COLLARED JAY (Cyanolyca viridicyanus)

Green Jay was one
of the species in the desert near Bagua Grande. This
sunrise preceded our search for Little Inca-Finch. The
Eastern Cordillera, our destination, is in the background.
(Photo by guide Richard Webster)
GREEN JAY (Cyanocorax yncas)
WHITE-TAILED JAY (Cyanocorax mystacalis)
Hirundinidae (Swallows)
BLUE-AND-WHITE SWALLOW (Pygochelidon cyanoleuca)
BROWN-BELLIED SWALLOW (Orochelidon murina)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)
GRAY-BREASTED MARTIN (Progne chalybea)
TUMBES SWALLOW (Tachycineta
stolzmanni)
BARN SWALLOW (Hirundo
rustica) [b]
Troglodytidae (Wrens)
FASCIATED WREN (Campylorhynchus
fasciatus)
THRUSH-LIKE WREN (Campylorhynchus turdinus)
GRAY-MANTLED WREN (Odontorchilus branickii)
SHARPE'S WREN (Cinnycerthia
olivascens) [*]
PERUVIAN WREN (Cinnycerthia
peruana) [*]
PLAIN-TAILED WREN (Pheugopedius euophrys schulenbergi)
[*]
SPECKLE-BREASTED WREN (SPECKLE-BREASTED)
(Pheugopedius sclateri paucimaculatus)
SPECKLE-BREASTED WREN (MARANON) (Pheugopedius sclateri sclateri)
SUPERCILIATED WREN (Cantorchilus superciliaris)
HOUSE WREN (Troglodytes
aedon)
MOUNTAIN WREN (Troglodytes
solstitialis)
SEDGE WREN (Cistothorus
platensis)
BAR-WINGED WOOD-WREN (Henicorhina leucoptera)
GRAY-BREASTED WOOD-WREN (Henicorhina leucophrys)
CHESTNUT-BREASTED WREN (Cyphorhinus thoracicus)
Polioptilidae (Gnatcatchers)
TROPICAL GNATCATCHER (WHITE-BROWED) (Polioptila plumbea bilineata)

Tropical
Gnatcatcher (Maranon) is one of many taxonomic puzzles
presented by populations in the Maranon separated from
those of the Pacific slope. (Photo by guide Richard
Webster)
TROPICAL
GNATCATCHER (MARANON) (Polioptila
plumbea maior)
Cinclidae (Dippers)
WHITE-CAPPED DIPPER (Cinclus leucocephalus)
Donacobiidae (Donacobius)
BLACK-CAPPED DONACOBIUS (Donacobius atricapilla)
Turdidae (Thrushes and Allies)
ANDEAN SOLITAIRE (Myadestes ralloides)
SWAINSON'S THRUSH (Catharus ustulatus)
[b]
PALE-EYED THRUSH (Turdus leucops)
PLUMBEOUS-BACKED THRUSH (Turdus reevei)
BLACK-BILLED THRUSH (Turdus ignobilis)
MARANON THRUSH (Turdus
maranonicus)
SLATY THRUSH (ANDEAN) (Turdus nigriceps nigriceps)
GREAT THRUSH (Turdus
fuscater)
CHIGUANCO THRUSH (Turdus chiguanco)
GLOSSY-BLACK THRUSH (Turdus serranus)
Mimidae (Mockingbirds and Thrashers)
LONG-TAILED MOCKINGBIRD (Mimus longicaudatus)
Motacillidae (Wagtails and Pipits)
PARAMO PIPIT (Anthus
bogotensis)
Parulidae (New World Warblers)
MASKED YELLOWTHROAT (MASKED) (Geothlypis aequinoctialis peruviana)
MASKED YELLOWTHROAT (BLACK-LORED) (Geothlypis aequinoctialis auricularis)
CERULEAN WARBLER (Setophaga cerulea)
[b]
TROPICAL PARULA (Setophaga pitiayumi)

Lunch at 2800m
means sun protection, from Joe's hat flaps to Mitch's
umbrella to several pairs of dark glasses (fight
catarcacts!), not to mention many flavors of SPF! (Photo
by guide Richard Webster)
BLACKBURNIAN
WARBLER (Setophaga fusca)
[b]
THREE-STRIPED WARBLER (Basileuterus tristriatus)
THREE-BANDED WARBLER (Basileuterus trifasciatus)
BLACK-CRESTED WARBLER (Myiothlypis nigrocristatus)
GRAY-AND-GOLD WARBLER (Myiothlypis fraseri)
RUSSET-CROWNED WARBLER (Myiothlypis coronatus)
CANADA WARBLER (Cardellina
canadensis) [b]
SLATE-THROATED REDSTART (Myioborus miniatus)
SPECTACLED REDSTART (Myioborus melanocephalus)
Coerebidae (Bananaquit)
BANANAQUIT (Coereba
flaveola)
Thraupidae (Tanagers and Allies)
MAGPIE TANAGER (Cissopis
leverianus)
WHITE-CAPPED TANAGER (Sericossypha albocristata)
RUFOUS-CRESTED TANAGER (Creurgops verticalis)
SUPERCILIARIED HEMISPINGUS (WHITE-BELLIED)
(Hemispingus superciliaris insignis)
OLEAGINOUS HEMISPINGUS (Hemispingus frontalis)
BLACK-EARED HEMISPINGUS (Hemispingus melanotis)
DRAB HEMISPINGUS (Hemispingus xanthophthalmus)
GRAY-HOODED BUSH TANAGER (Cnemoscopus rubrirostris)
RUFOUS-CHESTED TANAGER (Thlypopsis ornata)
BUFF-BELLIED TANAGER (Thlypopsis inornata)
CINEREOUS CONEBILL (Conirostrum cinereum)
BLUE-BACKED CONEBILL (Conirostrum sitticolor)
CAPPED CONEBILL (Conirostrum albifrons)
TIT-LIKE DACNIS (Xenodacnis parina)
YELLOW-CRESTED TANAGER (Tachyphonus rufiventer)
BLACK-BELLIED TANAGER (Ramphocelus melanogaster) [E]
SILVER-BEAKED TANAGER (Ramphocelus carbo)
BLUE-GRAY TANAGER (Thraupis episcopus)
PALM TANAGER (Thraupis
palmarum)
BLUE-CAPPED TANAGER (Thraupis cyanocephala)
BLUE-AND-YELLOW TANAGER (Thraupis bonariensis darwinii)
VERMILION TANAGER (Calochaetes coccineus)
HOODED MOUNTAIN-TANAGER (Buthraupis montana)
LACRIMOSE MOUNTAIN-TANAGER (Anisognathus lacrymosus)
SCARLET-BELLIED MOUNTAIN-TANAGER (Anisognathus igniventris)
GRASS-GREEN TANAGER (Chlorornis riefferii)
YELLOW-THROATED TANAGER (Iridosornis analis)

Yellow-scarfed
Tanager is one of the tanagers seen from the tower at
Owlet Lodge, the site of this sunset photo over the Andes.
(Photo by guide Richard Webster)
YELLOW-SCARFED
TANAGER (Iridosornis
reinhardti) [E]
ORANGE-EARED TANAGER (Chlorochrysa calliparaea)
TURQUOISE TANAGER (Tangara mexicana)
PARADISE TANAGER (Tangara chilensis)
GREEN-AND-GOLD TANAGER (Tangara schrankii)
GOLDEN TANAGER (Tangara
arthus)
SAFFRON-CROWNED TANAGER (Tangara xanthocephala)
GOLDEN-EARED TANAGER (Tangara chrysotis)
FLAME-FACED TANAGER (Tangara parzudakii)
YELLOW-BELLIED TANAGER (Tangara xanthogastra)
SPOTTED TANAGER (Tangara punctata)
BAY-HEADED TANAGER (Tangara gyrola)
GOLDEN-NAPED TANAGER (Tangara ruficervix)
METALLIC-GREEN TANAGER (Tangara labradorides)
BLUE-BROWED TANAGER (Tangara cyanotis)

Can one have too
many spatuletail pictures? Sure, but this one shows the
lovely gorget color a little better, along with the crown;
not quite up to tanager standards, but beautiful. (Photo
by guide Richard Webster)
BLUE-NECKED
TANAGER (Tangara cyanicollis)
BERYL-SPANGLED TANAGER (Tangara nigroviridis)
BLUE-AND-BLACK TANAGER (Tangara vassorii)
SILVERY TANAGER (Tangara viridicollis)
BLACK-FACED DACNIS (Dacnis lineata)
BLUE DACNIS (Dacnis
cayana)
PURPLE HONEYCREEPER (Cyanerpes caeruleus)
GOLDEN-COLLARED HONEYCREEPER (Iridophanes pulcherrimus)
SWALLOW TANAGER (Tersina viridis)
STREAKED SALTATOR (Saltator striatipectus immaculatus)
STREAKED SALTATOR (Saltator striatipectus peruvianus)
GRAYISH SALTATOR (Saltator coerulescens)
BLACK-COWLED SALTATOR (Saltator nigriceps)
GOLDEN-BILLED SALTATOR (Saltator aurantiirostris)
BUFF-THROATED SALTATOR (Saltator maximus)
SLATE-COLORED GROSBEAK (Saltator grossus) [*]
Emberizidae (Buntings, Sparrows and Allies)
PERUVIAN SIERRA-FINCH (Phrygilus punensis)
MOURNING SIERRA-FINCH (Phrygilus fruticeti)
PLUMBEOUS SIERRA-FINCH (Phrygilus unicolor)
ASH-BREASTED SIERRA-FINCH (Phrygilus plebejus)
BAND-TAILED SIERRA-FINCH (Phrygilus alaudinus)
CINEREOUS FINCH (Piezorhina cinerea) [E]
GRAY-WINGED INCA-FINCH (Incaspiza ortizi)
[E]

Buff-bridled
Inca-Finch was seen on the slopes of the Rio Maranon on
both sides of the river. (Photo by guide Richard Webster)
BUFF-BRIDLED
INCA-FINCH (Incaspiza laeta)
[E]
LITTLE INCA-FINCH (Incaspiza watkinsi)
[E]
PLAIN-TAILED WARBLING-FINCH (Poospiza alticola) [E]
COLLARED WARBLING-FINCH (Poospiza hispaniolensis)
BLUE-BLACK GRASSQUIT (Volatinia jacarina)
YELLOW-BELLIED SEEDEATER (Sporophila nigricollis)
PARROT-BILLED SEEDEATER (Sporophila peruviana)
DRAB SEEDEATER (Sporophila
simplex)

Chestnut-throated
Seedeater is one of a number of birds that have adapted to
rice fields that now occupy much of the lowlands. The
Cordillera de Colan stands in the background, one of the
homes of the Long-whiskered Owlet. (Photo by guide Richard
Webster)
CHESTNUT-THROATED
SEEDEATER (Sporophila telasco)
BAND-TAILED SEEDEATER (Catamenia analis)
PLAIN-COLORED SEEDEATER (Catamenia inornata)
DULL-COLORED GRASSQUIT (Tiaris obscurus)
MOUSTACHED FLOWERPIERCER (Diglossa mystacalis)
BLACK-THROATED FLOWERPIERCER (Diglossa brunneiventris)
WHITE-SIDED FLOWERPIERCER (Diglossa albilatera)

Leymebamba was a
base for visiting the high country, and also the site of
the wonderful museum about the Chachapoya. (Photo by guide
Richard Webster)
RUSTY
FLOWERPIERCER (Diglossa
sittoides)
DEEP-BLUE FLOWERPIERCER (Diglossa glauca)
BLUISH FLOWERPIERCER (Diglossa caerulescens)
MASKED FLOWERPIERCER (Diglossa cyanea)
SAFFRON FINCH (Sicalis
flaveola)
GRASSLAND YELLOW-FINCH (Sicalis luteola)
SULPHUR-THROATED FINCH (Sicalis taczanowskii)

Emerald-bellied
Puffleg was a daily visitor in small numbers to the
feeders at Owlet Lodge, and were on the "yardlists" of
many. (Photo by guide Richard Webster)
RED-CRESTED FINCH
(Coryphospingus cucullatus)
BLACK-CAPPED SPARROW (Arremon abeillei abeillei)
BLACK-CAPPED SPARROW (Arremon abeillei nigriceps)
GRAY-BROWED BRUSH-FINCH (Arremon assimilis)
YELLOW-BREASTED BRUSH-FINCH (Atlapetes latinuchus latinuchus)
YELLOW-BREASTED BRUSH-FINCH (Atlapetes latinuchus baroni)
RUFOUS-EARED BRUSH-FINCH (Atlapetes rufigenis) [E]
WHITE-WINGED BRUSH-FINCH (Atlapetes leucopterus)
WHITE-HEADED BRUSH-FINCH (Atlapetes albiceps)
BAY-CROWNED BRUSH-FINCH (Atlapetes seebohmi)
TUMBES SPARROW (Rhynchospiza
stolzmanni)
YELLOW-BROWED SPARROW (Ammodramus aurifrons)
RUFOUS-COLLARED SPARROW (Zonotrichia capensis)
COMMON BUSH-TANAGER (NORTHERN ANDES)
(Chlorospingus ophthalmicus hiaticolus)
ASHY-THROATED BUSH-TANAGER (Chlorospingus canigularis)
Cardinalidae (Cardinals and Allies)

Our hotel at
Banos del Inca was a delightful end to a trip that
included camping and some small-town hotels. (Photo by
guide Richard Webster)
HEPATIC TANAGER
(HIGHLAND) (Piranga flava
lutea)
WHITE-WINGED TANAGER (Piranga leucoptera)
RED-HOODED TANAGER (Piranga rubriceps)
GOLDEN-BELLIED GROSBEAK (Pheucticus chrysogaster)
Icteridae (Troupials and Allies)
PERUVIAN MEADOWLARK (Sturnella bellicosa)
SCRUB BLACKBIRD (Dives warszewiczi)
ORIOLE BLACKBIRD (Gymnomystax mexicanus)
SHINY COWBIRD (Molothrus
bonariensis)
GIANT COWBIRD (Molothrus
oryzivorus)
WHITE-EDGED ORIOLE (Icterus graceannae)
YELLOW-TAILED ORIOLE (Icterus mesomelas)
MOUNTAIN CACIQUE (GOLDEN-SHOULDERED)
(Cacicus chrysonotus peruvianus)
SCARLET-RUMPED CACIQUE (Cacicus uropygialis) [*]
YELLOW-RUMPED CACIQUE (Cacicus cela)
RUSSET-BACKED OROPENDOLA (Psarocolius angustifrons)
CRESTED OROPENDOLA (Psarocolius decumanus)
Fringillidae (Siskins, Crossbills, and
Allies)
PURPLE-THROATED EUPHONIA (Euphonia chlorotica)
THICK-BILLED EUPHONIA (Euphonia laniirostris)
GOLDEN-RUMPED EUPHONIA (Euphonia cyanocephala)
BRONZE-GREEN EUPHONIA (Euphonia mesochrysa)
ORANGE-BELLIED EUPHONIA (Euphonia xanthogaster)
BLUE-NAPED CHLOROPHONIA (Chlorophonia cyanea)
CHESTNUT-BREASTED CHLOROPHONIA (Chlorophonia pyrrhophrys)
LESSER GOLDFINCH (Spinus psaltria)
HOODED SISKIN (Spinus
magellanicus)

Yellow-tailed
Woolly Monkey is one of the rarest species we saw, and one
of the least expected. (Photo by guide Richard Webster)
YELLOW-TAILED
WOOLLY MONKEY (Lagothrix
flavicauda) [E]
GUAYAQUIL SQUIRREL (Sciurus stramineus)
NORTHERN MOUNTAIN VISCACHA (Lagidium peruanum)
SECHURAN FOX (Pseudalopex
sechurae)
TAYRA (Eira
barbara)
COLLARED PECCARY (Tayassu tajacu)
Opossum sp. by Virginia.
Bat sp., including drinking from the swimming pool at Jaen.
Lizards, including Blue-headed Whiptail, Dicrodon guttulatum,
in the coastal deserts, and Microlophus (formerly Tropidurus)
koepckeorum, Koepcke's Curly-tailed Lizard, on rocky slopes
above Balsas.
Many millipedes.
Incredible displays of moths around the lights at Owlet
Lodge.
A steady stream of lovely butterflies.
Totals for the tour: 548 bird taxa and 6 mammal taxa