For our tour description, itinerary, past triplists, dates, fees, and more, please VISIT OUR TOUR PAGE.
See this triplist in printable PDF format with media only on page 1.

A canopy lover, the Green-Shrike Vireo is notoriously difficult to see. We were so fortunate to see one up close where we could admire its beautiful, bright green plumage and serious, hook-tipped bill. (Photo by guide Lena Senko)
Since its inception in 1999, the Canopy Tower has gained worldwide fame for its proximity to some of the best birding hotspots in the country. Situated within the 55,000-acre Soberania National Park in central Panama, it is the ideal launching-off point for birders wishing to explore its diverse surroundings, which are home to some 525 species of birds and 105 species of mammals.
On this trip, we sampled a good chunk of this diversity and delighted in the common as well as the unexpected species that came our way. A beautiful Black Hawk-Eagle greeted us on our first day, soaring and calling right over the entrance road to the Canopy Tower -- a wonderful road for spotting White-whiskered and Black-chested puffbirds, Rufous and Broad-billed motmots, and a host of antbirds, antwrens, and woodcreepers. From the scenic upper story of the tower itself, we watched a slow, feeding Brown-throated Three-Toed Sloth while the treetops came alive with tanagers, tityras, warblers, and toucans.
A nearby birding hotspot, Ammo Pond, is a haven for Rufescent Tiger-Heron, Yellow-tailed Oriole, and Barred Antshrike. We admired these striking species while studying differences between the similar-looking Great Kiskadee, Boat-billed, Rusty-margined, and Social flycatchers. What's birding without a healthy flycatcher challenge, huh? We also observed two nests of Black-throated Mangos, one of which contained a couple of cute chicks. A visit to another birdy location, Summit Pond, yielded four species of kingfishers, including the teeny and tough-to-find American Pygmy Kingfisher. Nesting Boat-billed Herons were fun to watch as they clambered about on giant palm fronds, and we were in for quite a treat when we saw the rare and gorgeous Agami Heron and later Spectacled Owl!
A bird no one forgets on this tour is the dazzling male Blue Cotinga. Recently, a new observation tower was built near the start of the famous Pipeline Road, and it is the perfect perch from which to observe the canopy-loving cotinga, Collared Forest-Falcon, White-winged Becard, Yellow-backed Oriole, and Blue Dacnis, to name a few. The Pipeline Road itself is, of course, sensational. Many difficult goodies dwell along its track, including Spot-crowned Antshrike, Gray Elaenia, Great Tinamou, and four trogon species. Great Jacamar, Black-striped Woodcreeper, Speckled Mourner, and Streak-chested Antpitta were nice finds, too.
Other highlights included a Common Potoo on a day roost and a night drive where we saw not one, not two, but five Kinkajous, along with opossums, a Common Pauraque, and even a surprise Crested Owl (that was something!). Aside from the birds, there were many other enjoyments to be had, such as the cool, refreshing breeze up on the Canopy Tower and the sun-ripened sweetness of abundant papayas, melons, and pineapples. A trip to central Panama would not be complete without a visit to the Panama Canal, where we absorbed the history of this monumental human achievement and watched a giant cargo ship pass through the locks.
We owe heaping thanks to the Canopy Tower staff for their fantastic hospitality and to our incredibly keen local guide, Alexis Sanchez. Thanks also to each one of you for your lively energy and enthusiasm…John and I look forward to seeing you again down the road!
In the words of the Whooping Motmot…
"Whoop-whoop!"
--Lena
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Tinamidae (Tinamous)
GREAT TINAMOU (Tinamus major)
Anatidae (Ducks, Geese, and Waterfowl)
BLACK-BELLIED WHISTLING-DUCK (Dendrocygna autumnalis)
MUSCOVY DUCK (Cairina moschata)
Cracidae (Guans, Chachalacas, and Curassows)
GRAY-HEADED CHACHALACA (Ortalis cinereiceps)
Fregatidae (Frigatebirds)

A Common Potoo sits perfectly camouflaged on its day perch. (Photo by guide Lena Senko)
MAGNIFICENT FRIGATEBIRD (Fregata magnificens) Anhingidae (Anhingas)
ANHINGA (Anhinga anhinga)
Pelecanidae (Pelicans)
BROWN PELICAN (Pelecanus occidentalis)
Ardeidae (Herons, Egrets, and Bitterns)
RUFESCENT TIGER-HERON (Tigrisoma lineatum)
GREAT EGRET (Ardea alba)
SNOWY EGRET (Egretta thula)
LITTLE BLUE HERON (Egretta caerulea)
CATTLE EGRET (Bubulcus ibis)
GREEN HERON (Butorides virescens)
STRIATED HERON (Butorides striata)
AGAMI HERON (Agamia agami)
BOAT-BILLED HERON (SOUTHERN) (Cochlearius cochlearius panamensis) [N]
Threskiornithidae (Ibises and Spoonbills)
WHITE IBIS (Eudocimus albus)
Cathartidae (New World Vultures)
BLACK VULTURE (Coragyps atratus)
TURKEY VULTURE (Cathartes aura)
KING VULTURE (Sarcoramphus papa)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus)
Accipitridae (Hawks, Eagles, and Kites)
GRAY-HEADED KITE (Leptodon cayanensis) [*]
BLACK HAWK-EAGLE (Spizaetus tyrannus)
DOUBLE-TOOTHED KITE (Harpagus bidentatus)
CRANE HAWK (Geranospiza caerulescens)
GRAY-LINED HAWK (Buteo nitidus)
SHORT-TAILED HAWK (Buteo brachyurus)
Rallidae (Rails, Gallinules, and Coots)
WHITE-THROATED CRAKE (Laterallus albigularis) [*]
GRAY-NECKED WOOD-RAIL (Aramides cajaneus) [*]
COMMON GALLINULE (Gallinula galeata)
Jacanidae (Jacanas)
WATTLED JACANA (Jacana jacana)
Scolopacidae (Sandpipers and Allies)
SPOTTED SANDPIPER (Actitis macularius)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia)
PALE-VENTED PIGEON (Patagioenas cayennensis)
SCALED PIGEON (Patagioenas speciosa)
SHORT-BILLED PIGEON (Patagioenas nigrirostris)
RUDDY GROUND-DOVE (Columbina talpacoti)
WHITE-TIPPED DOVE (Leptotila verreauxi)
GRAY-CHESTED DOVE (Leptotila cassini)
Cuculidae (Cuckoos)

Finding owls by night is fantastic. Spotting an owl during the day is even more special. This Spectacled Owl was discovered on its day roost, peeping at us from behind a leaf. (Photo by guide Lena Senko)
SQUIRREL CUCKOO (Piaya cayana) GREATER ANI (Crotophaga major)
SMOOTH-BILLED ANI (Crotophaga ani)
Strigidae (Owls)
CRESTED OWL (Lophostrix cristata)
SPECTACLED OWL (Pulsatrix perspicillata)
Caprimulgidae (Nightjars and Allies)
COMMON NIGHTHAWK (Chordeiles minor)
COMMON PAURAQUE (Nyctidromus albicollis)
Nyctibiidae (Potoos)
COMMON POTOO (Nyctibius griseus)
Apodidae (Swifts)
SHORT-TAILED SWIFT (Chaetura brachyura)
Trochilidae (Hummingbirds)
WHITE-NECKED JACOBIN (Florisuga mellivora)
BAND-TAILED BARBTHROAT (Threnetes ruckeri)
LONG-BILLED HERMIT (Phaethornis longirostris)
STRIPE-THROATED HERMIT (Phaethornis striigularis)
PURPLE-CROWNED FAIRY (Heliothryx barroti)
BLACK-THROATED MANGO (Anthracothorax nigricollis) [N]
BLUE-CHESTED HUMMINGBIRD (Amazilia amabilis)
SNOWY-BELLIED HUMMINGBIRD (Amazilia edward edward)
RUFOUS-TAILED HUMMINGBIRD (Amazilia tzacatl)
VIOLET-BELLIED HUMMINGBIRD (Damophila julie)
Trogonidae (Trogons)
SLATY-TAILED TROGON (Trogon massena)
WHITE-TAILED TROGON (Trogon chionurus)
GARTERED TROGON (Trogon caligatus) [*]
BLACK-THROATED TROGON (Trogon rufus)
Momotidae (Motmots)
WHOOPING MOTMOT (WHOOPING) (Momotus subrufescens conexus)
RUFOUS MOTMOT (Baryphthengus martii)
BROAD-BILLED MOTMOT (Electron platyrhynchum)
Alcedinidae (Kingfishers)
RINGED KINGFISHER (Megaceryle torquata)
AMAZON KINGFISHER (Chloroceryle amazona)
GREEN KINGFISHER (Chloroceryle americana)
AMERICAN PYGMY KINGFISHER (Chloroceryle aenea)
Bucconidae (Puffbirds)
WHITE-NECKED PUFFBIRD (Notharchus hyperrhynchus)

Sleepy as can be, a Brown-throated Three-Toed Sloth takes a snooze in a Ceiba tree at Metro Park. (Photo by guide Lena Senko)
BLACK-BREASTED PUFFBIRD (Notharchus pectoralis) WHITE-WHISKERED PUFFBIRD (Malacoptila panamensis)
Galbulidae (Jacamars)
GREAT JACAMAR (Jacamerops aureus)
Ramphastidae (Toucans)
COLLARED ARACARI (Pteroglossus torquatus)
BLACK-MANDIBLED TOUCAN (CHESTNUT-MANDIBLED) (Ramphastos ambiguus swainsonii)
KEEL-BILLED TOUCAN (Ramphastos sulfuratus)
Picidae (Woodpeckers)
BLACK-CHEEKED WOODPECKER (Melanerpes pucherani)
RED-CROWNED WOODPECKER (Melanerpes rubricapillus)
CINNAMON WOODPECKER (Celeus loricatus)
LINEATED WOODPECKER (Dryocopus lineatus)
CRIMSON-CRESTED WOODPECKER (Campephilus melanoleucos)
Falconidae (Falcons and Caracaras)
SLATY-BACKED FOREST-FALCON (Micrastur mirandollei) [*]
COLLARED FOREST-FALCON (Micrastur semitorquatus)
YELLOW-HEADED CARACARA (Milvago chimachima)
LAUGHING FALCON (Herpetotheres cachinnans) [*]
Psittacidae (Parrots)
ORANGE-CHINNED PARAKEET (Brotogeris jugularis)
BROWN-HOODED PARROT (Pyrilia haematotis) [*]
BLUE-HEADED PARROT (Pionus menstruus)
MEALY PARROT (Amazona farinosa)
Thamnophilidae (Typical Antbirds)
FASCIATED ANTSHRIKE (Cymbilaimus lineatus)
BARRED ANTSHRIKE (Thamnophilus doliatus)
BLACK-CROWNED ANTSHRIKE (Thamnophilus atrinucha)
SPOT-CROWNED ANTVIREO (Dysithamnus puncticeps)
CHECKER-THROATED ANTWREN (Epinecrophylla fulviventris)

Numerous species of butterflies delighted us as well, like this Heliconia. (Photo by guide Lena Senko)
MOUSTACHED ANTWREN (Myrmotherula ignota) WHITE-FLANKED ANTWREN (Myrmotherula axillaris)
DOT-WINGED ANTWREN (Microrhopias quixensis)
DUSKY ANTBIRD (Cercomacra tyrannina)
WHITE-BELLIED ANTBIRD (Myrmeciza longipes)
CHESTNUT-BACKED ANTBIRD (Myrmeciza exsul)
BICOLORED ANTBIRD (Gymnopithys leucaspis bicolor)
SPOTTED ANTBIRD (Hylophylax naevioides)
Grallariidae (Antpittas)
STREAK-CHESTED ANTPITTA (Hylopezus perspicillatus)
Formicariidae (Antthrushes)
BLACK-FACED ANTTHRUSH (Formicarius analis)
Furnariidae (Ovenbirds and Woodcreepers)
OLIVACEOUS WOODCREEPER (GRAYISH) (Sittasomus griseicapillus sylvioides)
PLAIN-BROWN WOODCREEPER (Dendrocincla fuliginosa)
NORTHERN BARRED-WOODCREEPER (Dendrocolaptes sanctithomae)
COCOA WOODCREEPER (Xiphorhynchus susurrans)
BLACK-STRIPED WOODCREEPER (Xiphorhynchus lachrymosus)
PLAIN XENOPS (Xenops minutus)
Tyrannidae (Tyrant Flycatchers)
BROWN-CAPPED TYRANNULET (Ornithion brunneicapillus)
SOUTHERN BEARDLESS-TYRANNULET (Camptostoma obsoletum)
YELLOW TYRANNULET (Capsiempis flaveola)
YELLOW-CROWNED TYRANNULET (Tyrannulus elatus)
FOREST ELAENIA (Myiopagis gaimardii)
GRAY ELAENIA (Myiopagis caniceps absita)

A female Spotted Antbird perches in the undergrowth. (Photo by guide Lena Senko)
GREENISH ELAENIA (Myiopagis viridicata) [*] YELLOW-BELLIED ELAENIA (Elaenia flavogaster) [*]
OCHRE-BELLIED FLYCATCHER (Mionectes oleagineus)
PALTRY TYRANNULET (Zimmerius vilissimus)
BLACK-CAPPED PYGMY-TYRANT (Myiornis atricapillus)
SOUTHERN BENTBILL (Oncostoma olivaceum)
COMMON TODY-FLYCATCHER (Todirostrum cinereum)
BLACK-HEADED TODY-FLYCATCHER (Todirostrum nigriceps)
BROWNISH TWISTWING (Cnipodectes subbrunneus) [*]
OLIVACEOUS FLATBILL (Rhynchocyclus olivaceus)
YELLOW-OLIVE FLYCATCHER (YELLOW-OLIVE) (Tolmomyias sulphurescens flavoolivaceus)
YELLOW-MARGINED FLYCATCHER (YELLOW-MARGINED) (Tolmomyias assimilis flavotectus)
BLACK-TAILED FLYCATCHER (Myiobius atricaudus)
ACADIAN FLYCATCHER (Empidonax virescens) [*]
BRIGHT-RUMPED ATTILA (Attila spadiceus)
RUFOUS MOURNER (Rhytipterna holerythra)
DUSKY-CAPPED FLYCATCHER (Myiarchus tuberculifer)
PANAMA FLYCATCHER (Myiarchus panamensis)
GREAT CRESTED FLYCATCHER (Myiarchus crinitus)
LESSER KISKADEE (Pitangus lictor)
GREAT KISKADEE (Pitangus sulphuratus)
BOAT-BILLED FLYCATCHER (Megarynchus pitangua)
RUSTY-MARGINED FLYCATCHER (Myiozetetes cayanensis)
SOCIAL FLYCATCHER (Myiozetetes similis)
STREAKED FLYCATCHER (Myiodynastes maculatus)
PIRATIC FLYCATCHER (Legatus leucophaius)
TROPICAL KINGBIRD (Tyrannus melancholicus)
FORK-TAILED FLYCATCHER (Tyrannus savana)
Cotingidae (Cotingas)
PURPLE-THROATED FRUITCROW (Querula purpurata)
BLUE COTINGA (Cotinga nattererii)
Pipridae (Manakins)
LANCE-TAILED MANAKIN (Chiroxiphia lanceolata)
RED-CAPPED MANAKIN (Ceratopipra mentalis)
GOLDEN-COLLARED MANAKIN (Manacus vitellinus)
BLUE-CROWNED MANAKIN (Lepidothrix coronata)
Tityridae (Tityras and Allies)

This sleek-looking Streaked Flycatcher was a nice find at Ammo Pond. (Photo by guide Lena Senko)
BLACK-CROWNED TITYRA (Tityra inquisitor) MASKED TITYRA (Tityra semifasciata)
RUSSET-WINGED SCHIFFORNIS (Schiffornis stenorhyncha panamensis)
SPECKLED MOURNER (Laniocera rufescens)
CINNAMON BECARD (Pachyramphus cinnamomeus) [N]
WHITE-WINGED BECARD (Pachyramphus polychopterus)
Vireonidae (Vireos)
YELLOW-THROATED VIREO (Vireo flavifrons)
YELLOW-GREEN VIREO (Vireo flavoviridis)
SCRUB GREENLET (Hylophilus flavipes)
GOLDEN-FRONTED GREENLET (Hylophilus aurantiifrons)
LESSER GREENLET (Hylophilus decurtatus)
GREEN SHRIKE-VIREO (Vireolanius pulchellus)
Corvidae (Crows, Jays, and Magpies)
BLACK-CHESTED JAY (Cyanocorax affinis) [*]
Hirundinidae (Swallows)
SOUTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx ruficollis)
GRAY-BREASTED MARTIN (Progne chalybea)
MANGROVE SWALLOW (Tachycineta albilinea)
Troglodytidae (Wrens)
HOUSE WREN (Troglodytes aedon)
BLACK-BELLIED WREN (Pheugopedius fasciatoventris)
RUFOUS-BREASTED WREN (Pheugopedius rutilus)
RUFOUS-AND-WHITE WREN (Thryophilus rufalbus)
PLAIN WREN (Cantorchilus modestus)
BAY WREN (Cantorchilus nigricapillus) [*]
BUFF-BREASTED WREN (Cantorchilus leucotis)
WHITE-BREASTED WOOD-WREN (Henicorhina leucosticta)
SONG WREN (Cyphorhinus phaeocephalus)
Polioptilidae (Gnatcatchers)
LONG-BILLED GNATWREN (Ramphocaenus melanurus)
TROPICAL GNATCATCHER (Polioptila plumbea)
Turdidae (Thrushes and Allies)
CLAY-COLORED THRUSH (Turdus grayi)
Mimidae (Mockingbirds and Thrashers)
TROPICAL MOCKINGBIRD (Mimus gilvus)
Parulidae (New World Warblers)

A Red-naped Tamarin sits and ponders the meaning of life. (Photo by guide Lena Senko)
NORTHERN WATERTHRUSH (Parkesia noveboracensis) GOLDEN-WINGED WARBLER (Vermivora chrysoptera)
BLACK-AND-WHITE WARBLER (Mniotilta varia)
PROTHONOTARY WARBLER (Protonotaria citrea)
TENNESSEE WARBLER (Oreothlypis peregrina)
MOURNING WARBLER (Geothlypis philadelphia)
BAY-BREASTED WARBLER (Setophaga castanea)
YELLOW WARBLER (Setophaga petechia)
CHESTNUT-SIDED WARBLER (Setophaga pensylvanica)
RUFOUS-CAPPED WARBLER (Basileuterus rufifrons)
Thraupidae (Tanagers and Allies)
WHITE-SHOULDERED TANAGER (Tachyphonus luctuosus)
CRIMSON-BACKED TANAGER (Ramphocelus dimidiatus)
FLAME-RUMPED TANAGER (Ramphocelus flammigerus)
BLUE-GRAY TANAGER (Thraupis episcopus)
PALM TANAGER (Thraupis palmarum)
GOLDEN-HOODED TANAGER (Tangara larvata)
PLAIN-COLORED TANAGER (Tangara inornata)
BLUE DACNIS (Dacnis cayana)
RED-LEGGED HONEYCREEPER (Cyanerpes cyaneus)
GREEN HONEYCREEPER (Chlorophanes spiza)
VARIABLE SEEDEATER (Sporophila corvina hoffmannii)
YELLOW-BELLIED SEEDEATER (Sporophila nigricollis)
ROSY THRUSH-TANAGER (Rhodinocichla rosea)
STREAKED SALTATOR (Saltator striatipectus)
BUFF-THROATED SALTATOR (Saltator maximus)
SLATE-COLORED GROSBEAK (Saltator grossus) [*]
Emberizidae (Buntings and New World Sparrows)
ORANGE-BILLED SPARROW (Arremon aurantiirostris)
BLACK-STRIPED SPARROW (Arremonops conirostris)
Cardinalidae (Cardinals and Allies)

All loaded up and ready to go in the Birdmobile. (Photo by guide Lena Senko)
SUMMER TANAGER (Piranga rubra) RED-CROWNED ANT-TANAGER (Habia rubica)
RED-THROATED ANT-TANAGER (Habia fuscicauda)
BLUE-BLACK GROSBEAK (Cyanocompsa cyanoides)
Icteridae (Troupials and Allies)
RED-BREASTED BLACKBIRD (Sturnella militaris)
GREAT-TAILED GRACKLE (Quiscalus mexicanus)
GIANT COWBIRD (Molothrus oryzivorus)
ORCHARD ORIOLE (Icterus spurius)
YELLOW-BACKED ORIOLE (Icterus chrysater)
YELLOW-TAILED ORIOLE (Icterus mesomelas)
BALTIMORE ORIOLE (Icterus galbula)
SCARLET-RUMPED CACIQUE (SCARLET-RUMPED) (Cacicus uropygialis microrhynchus)
YELLOW-RUMPED CACIQUE (Cacicus cela)
CHESTNUT-HEADED OROPENDOLA (Psarocolius wagleri)
Fringillidae (Siskins, Crossbills, and Allies)
YELLOW-CROWNED EUPHONIA (Euphonia luteicapilla)
THICK-BILLED EUPHONIA (Euphonia laniirostris)
FULVOUS-VENTED EUPHONIA (Euphonia fulvicrissa)
COMMON OPOSSUM (Didelphis marsupialis)
CENTRAL AMERICAN WOOLY OPOSSUM (Caluromys derbianus)
COMMON TENT-MAKING BAT (Uroderma bilobatum)
RED-NAPED TAMARIN (Saguinus geoffroyi)
MANTLED HOWLER MONKEY (Alouatta palliata)
WHITE-THROATED CAPUCHIN (Cebus capucinus)
HOFFMANN'S TWO-TOED SLOTH (Choloepus hoffmanni)
BROWN-THROATED THREE-TOED SLOTH (Bradypus variegatus)
NORTHERN TAMANDUA (Tamandua mexicana)
VARIEGATED SQUIRREL (Sciurus variegatoides)
RED-TAILED SQUIRREL (Sciurus granatensis)
CENTRAL AMERICAN AGOUTI (Dasyprocta punctata)
WHITE-NOSED COATI (Nasua narica)
KINKAJOU (Potos flavus)
MAMMAL ADDITION:
Pallas's long-tongued bat (Glossophaga soricina) -- According to Alexis, this is the common bat species seen flitting about in the Canopy Tower. It is also the one we saw hanging from the tip of a branch at Summit Pond.
REPTILES & AMPHIBIANS
Canopy Lizard (Polychrus) -- A.k.a. Berthold's Bush Anole. Seen, appropriately, in the canopy on the first day. An exciting find, since this long-tailed species is uncommon and is hardly ever seen due to its height-loving habits.
Green Iguana (Iguana iguana) -- Seen lounging on the limbs at Ammo Pond.
Common Basilisk (Basiliscus basiliscus) -- Colloquially called the Jesus Christ Lizard for its ability to run on the surface of the water. We saw a nice male on the grounds of the Gamboa Rainforest Resort.
Leaf Toad species -- There are numerous species of these small toads (leaf toads) in the Neotropics. They all have in common the superficial resemblance to a dead leaf on the ground, often with a white streak down the back.
BUTTERFLIES: There are hundreds of species in the vicinity of Canopy Tower. Our attention was mainly focused on the birds, hence this short list. Here are a few that were identified with certainty:
Many-banded Daggerwing (Marpesia chiron)
Banded Peacock (Papilio crino) -- Extremely common.
Orion Cecropian (Historis odius) -- A type of Leafwing butterfly.
Cramer’s Scarlet-eye (Nascus broteas) -- Female on the window at the Canopy Tower.
Totals for the tour: 224 bird taxa and 14 mammal taxa