Pennsylvania's Warblers & More: from Cerulean Warbler to Henslow's Sparrow 2019
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See this triplist in printable PDF format with media only on page 1.

Though we were initially deterred by passing thunderstorms, a bit of patience led to a memorable sighting of this Henslow's Sparrow near Punxsutawney, home of Groundhog Day! Photo by leader Tom Johnson.
The third annual spring Pennsylvania tour underscored the impressive diversity of breeding songbirds in this beautiful, largely forested state. Our group birded from the banks of the lower Susquehanna River to the ridges of the Appalachian Mountains to the grasslands near Punxsutawney and the varied forests of the state's northern tier. We found 29 species of wood-warblers and were fortunate to see many of these fine birds singing and engaged in other breeding behavior.
Some highlights of the adventure included Prothonotary Warblers, Cerulean Warbler, Golden-winged Warblers, Canada Warblers, Mourning Warbler, and Kentucky Warbler. Oh, but wait - we saw more than "just" warblers: Sandhill Cranes, Henslow's Sparrow, Willow and Alder flycatchers, Brown Thrasher, and Orchard Oriole come to mind, too! For more details on particular species, check out the annotated species list below.
Thanks to our excellent group for your enthusiastic collaboration in the field during this exploration of the Keystone State!
-Tom
KEYS FOR THIS LIST
One of the following keys may be shown in brackets for individual species as appropriate: * = heard only, I = introduced, E = endemic, N = nesting, a = austral migrant, b = boreal migrant
Anatidae (Ducks, Geese, and Waterfowl)
CANADA GOOSE (Branta canadensis)
WOOD DUCK (Aix sponsa)
MALLARD (Anas platyrhynchos)

In the mixed forests of Bear Meadows, we visited with gems such as this male Magnolia Warbler. Photo by group member Kim Nelson.
BUFFLEHEAD (Bucephala albeola)
HOODED MERGANSER (Lophodytes cucullatus)
COMMON MERGANSER (Mergus merganser)
RED-BREASTED MERGANSER (Mergus serrator)
Phasianidae (Pheasants, Grouse, and Allies)
WILD TURKEY (Meleagris gallopavo)
Columbidae (Pigeons and Doves)
ROCK PIGEON (Columba livia) [I]
MOURNING DOVE (Zenaida macroura)
Cuculidae (Cuckoos)
YELLOW-BILLED CUCKOO (Coccyzus americanus)
BLACK-BILLED CUCKOO (Coccyzus erythropthalmus)
Caprimulgidae (Nightjars and Allies)
COMMON NIGHTHAWK (Chordeiles minor)
EASTERN WHIP-POOR-WILL (Antrostomus vociferus)
Apodidae (Swifts)
CHIMNEY SWIFT (Chaetura pelagica)
Trochilidae (Hummingbirds)
RUBY-THROATED HUMMINGBIRD (Archilochus colubris)
Rallidae (Rails, Gallinules, and Coots)
VIRGINIA RAIL (Rallus limicola)
SORA (Porzana carolina) [*]
Charadriidae (Plovers and Lapwings)
KILLDEER (Charadrius vociferus)

One of the highlights of our night birding excursion at the Scotia Barrens was this handsome American Woodcock. This bizarre and amazing shorebird froze in the headlights of our van and walked around with his curious, bobbing gait for several minutes before returning to display from the brush. Photo by leader Tom Johnson.
Scolopacidae (Sandpipers and Allies)
AMERICAN WOODCOCK (Scolopax minor)
SPOTTED SANDPIPER (Actitis macularius)
Phalacrocoracidae (Cormorants and Shags)
DOUBLE-CRESTED CORMORANT (Phalacrocorax auritus)
Ardeidae (Herons, Egrets, and Bitterns)
GREAT BLUE HERON (Ardea herodias)
GREAT EGRET (Ardea alba)
GREEN HERON (Butorides virescens)
YELLOW-CROWNED NIGHT-HERON (Nyctanassa violacea)
Cathartidae (New World Vultures)
TURKEY VULTURE (Cathartes aura)
Pandionidae (Osprey)
OSPREY (Pandion haliaetus)
Accipitridae (Hawks, Eagles, and Kites)
SHARP-SHINNED HAWK (Accipiter striatus)
BALD EAGLE (Haliaeetus leucocephalus)

After we heard the loud "Three-Eight!" song of this Yellow-throated Vireo, it took a few minutes to actually spot the bird since it was singing from its nest! Photo by group member Nick Winograd.
BROAD-WINGED HAWK (Buteo platypterus)
RED-TAILED HAWK (Buteo jamaicensis)
Strigidae (Owls)
BARRED OWL (Strix varia) [*]
Alcedinidae (Kingfishers)
BELTED KINGFISHER (Megaceryle alcyon)
Picidae (Woodpeckers)
YELLOW-BELLIED SAPSUCKER (Sphyrapicus varius)
RED-HEADED WOODPECKER (Melanerpes erythrocephalus)
RED-BELLIED WOODPECKER (Melanerpes carolinus)
DOWNY WOODPECKER (Dryobates pubescens)
HAIRY WOODPECKER (Dryobates villosus)

Our eyes were regularly seared by close views of colorful warblers such as this male Blackburnian Warbler. Photo by leader Tom Johnson.
PILEATED WOODPECKER (Dryocopus pileatus)
NORTHERN FLICKER (YELLOW-SHAFTED) (Colaptes auratus auratus)
Falconidae (Falcons and Caracaras)
AMERICAN KESTREL (Falco sparverius)
PEREGRINE FALCON (Falco peregrinus)
Tyrannidae (Tyrant Flycatchers)
EASTERN WOOD-PEWEE (Contopus virens)
ACADIAN FLYCATCHER (Empidonax virescens)
ALDER FLYCATCHER (Empidonax alnorum)
WILLOW FLYCATCHER (Empidonax traillii)

A highly skilled Belted Kingfisher was busy subduing this Brook Trout when we spotted it. Photo by group member Kim Nelson.
LEAST FLYCATCHER (Empidonax minimus)
EASTERN PHOEBE (Sayornis phoebe)
GREAT CRESTED FLYCATCHER (Myiarchus crinitus)
EASTERN KINGBIRD (Tyrannus tyrannus)
Vireonidae (Vireos, Shrike-Babblers, and Erpornis)
YELLOW-THROATED VIREO (Vireo flavifrons)
BLUE-HEADED VIREO (Vireo solitarius)
WARBLING VIREO (Vireo gilvus)
RED-EYED VIREO (Vireo olivaceus)
Corvidae (Crows, Jays, and Magpies)
BLUE JAY (Cyanocitta cristata)
AMERICAN CROW (Corvus brachyrhynchos)

A male Canada Warbler stared at us as we passed through his nesting territory along the Pine Creek in Tioga County. Photo by leader Tom Johnson.
FISH CROW (Corvus ossifragus)
COMMON RAVEN (Corvus corax)
Alaudidae (Larks)
HORNED LARK (Eremophila alpestris)
Hirundinidae (Swallows)
NORTHERN ROUGH-WINGED SWALLOW (Stelgidopteryx serripennis)
TREE SWALLOW (Tachycineta bicolor)
BARN SWALLOW (Hirundo rustica)
CLIFF SWALLOW (Petrochelidon pyrrhonota)
Paridae (Tits, Chickadees, and Titmice)
CAROLINA CHICKADEE (Poecile carolinensis)
BLACK-CAPPED CHICKADEE (Poecile atricapillus)
TUFTED TITMOUSE (Baeolophus bicolor)
Sittidae (Nuthatches)
WHITE-BREASTED NUTHATCH (Sitta carolinensis)
Certhiidae (Treecreepers)
BROWN CREEPER (Certhia americana)
Troglodytidae (Wrens)
HOUSE WREN (Troglodytes aedon)

A male Mourning Warbler offered his "churry-churry-churry-churry-churry-o" proclamations from a treefall gap near Colton Point State Park in Tioga County. This scarce warbler was a nice highlight for many in the group. Photo by group member Nick Winograd.
MARSH WREN (Cistothorus palustris)
CAROLINA WREN (Thryothorus ludovicianus)
Polioptilidae (Gnatcatchers)
BLUE-GRAY GNATCATCHER (Polioptila caerulea)
Regulidae (Kinglets)
GOLDEN-CROWNED KINGLET (Regulus satrapa)
Turdidae (Thrushes and Allies)
EASTERN BLUEBIRD (Sialia sialis)
VEERY (Catharus fuscescens)
SWAINSON'S THRUSH (Catharus ustulatus)
HERMIT THRUSH (Catharus guttatus)
WOOD THRUSH (Hylocichla mustelina)
AMERICAN ROBIN (Turdus migratorius)
Mimidae (Mockingbirds and Thrashers)
GRAY CATBIRD (Dumetella carolinensis)
BROWN THRASHER (Toxostoma rufum)
NORTHERN MOCKINGBIRD (Mimus polyglottos)
Sturnidae (Starlings)
EUROPEAN STARLING (Sturnus vulgaris)
Bombycillidae (Waxwings)
CEDAR WAXWING (Bombycilla cedrorum)
Fringillidae (Finches, Euphonias, and Allies)
HOUSE FINCH (Haemorhous mexicanus)
AMERICAN GOLDFINCH (Spinus tristis)
Passerellidae (New World Sparrows)
GRASSHOPPER SPARROW (Ammodramus savannarum)
CHIPPING SPARROW (Spizella passerina)
FIELD SPARROW (Spizella pusilla)
DARK-EYED JUNCO (SLATE-COLORED) (Junco hyemalis carolinensis)

Grasshopper Sparrows were some of the many fine grassland birds we studied near Punxsutawney. Photo by group member Kim Nelson.
WHITE-THROATED SPARROW (Zonotrichia albicollis)
HENSLOW'S SPARROW (Centronyx henslowii)
SONG SPARROW (Melospiza melodia)
SWAMP SPARROW (Melospiza georgiana)
EASTERN TOWHEE (Pipilo erythrophthalmus)
Icteriidae (Yellow-breasted Chat)
YELLOW-BREASTED CHAT (Icteria virens)
Icteridae (Troupials and Allies)
BOBOLINK (Dolichonyx oryzivorus)
EASTERN MEADOWLARK (Sturnella magna)
ORCHARD ORIOLE (Icterus spurius)
BALTIMORE ORIOLE (Icterus galbula)
RED-WINGED BLACKBIRD (Agelaius phoeniceus)
BROWN-HEADED COWBIRD (Molothrus ater)
COMMON GRACKLE (Quiscalus quiscula)
Parulidae (New World Warblers)
OVENBIRD (Seiurus aurocapilla)
WORM-EATING WARBLER (Helmitheros vermivorum)
LOUISIANA WATERTHRUSH (Parkesia motacilla)
NORTHERN WATERTHRUSH (Parkesia noveboracensis) [*]

This Eastern Box Turtle was crossing a busy road in Lebanon County, so we stopped, examined it for a minute, and then carefully placed it off to the side of the road in its original direction of travel. Vehicle traffic is tough on reptiles, so it's a good idea to help them whenever you get the chance! Photo by group member Kim Nelson.
GOLDEN-WINGED WARBLER (Vermivora chrysoptera)
BLUE-WINGED WARBLER (Vermivora cyanoptera)
BLACK-AND-WHITE WARBLER (Mniotilta varia)
PROTHONOTARY WARBLER (Protonotaria citrea)
TENNESSEE WARBLER (Oreothlypis peregrina)
NASHVILLE WARBLER (RUFICAPILLA) (Oreothlypis ruficapilla ruficapilla) [*]
MOURNING WARBLER (Geothlypis philadelphia)
KENTUCKY WARBLER (Geothlypis formosa)
COMMON YELLOWTHROAT (Geothlypis trichas)
HOODED WARBLER (Setophaga citrina)
AMERICAN REDSTART (Setophaga ruticilla)
CERULEAN WARBLER (Setophaga cerulea)
NORTHERN PARULA (Setophaga americana)
MAGNOLIA WARBLER (Setophaga magnolia)
BAY-BREASTED WARBLER (Setophaga castanea)
BLACKBURNIAN WARBLER (Setophaga fusca)
YELLOW WARBLER (Setophaga petechia)
CHESTNUT-SIDED WARBLER (Setophaga pensylvanica)
BLACKPOLL WARBLER (Setophaga striata)
BLACK-THROATED BLUE WARBLER (Setophaga caerulescens cairnsi)
PINE WARBLER (Setophaga pinus)

Common Yellowthroat is one of the most abundant species of warblers that we encountered. These familiar beauties are frequently found along field edges and near wetlands. Photo by group member Nick Winograd.
YELLOW-THROATED WARBLER (ALBILORA) (Setophaga dominica albilora)
PRAIRIE WARBLER (Setophaga discolor)
BLACK-THROATED GREEN WARBLER (Setophaga virens)
CANADA WARBLER (Cardellina canadensis)
Cardinalidae (Cardinals and Allies)
SCARLET TANAGER (Piranga olivacea)
NORTHERN CARDINAL (Cardinalis cardinalis)
ROSE-BREASTED GROSBEAK (Pheucticus ludovicianus)
INDIGO BUNTING (Passerina cyanea)
Passeridae (Old World Sparrows)
HOUSE SPARROW (Passer domesticus)
VIRGINIA OPOSSUM (Didelphis virginianus)
EASTERN COTTONTAIL (Sylvilagus floridanus)
PLAIN EASTERN CHIPMUNK (Tamias striatus)
WOODCHUCK (Marmota monax)
EASTERN GRAY SQUIRREL (Sciurus carolinensis)
FOX SQUIRREL (Sciurus niger)
MUSKRAT (Ondatra zibethica)
WHITE-TAILED DEER (Odocoileus virginianus)
Totals for the tour: 138 bird taxa and 8 mammal taxa